A revision of Biting Midges of the Subgenera Forcipomyia (Metaforcipomyia) and F. (Saliohelea) from Costa Rica (Diptera: Ceratopogonidae) Author Spinelli, Gustavo R. Author Marino, Pablo I. Author Borkent, Art text Zootaxa 2012 3419 1 52 journal article 10.5281/zenodo.215031 215f451e-15f2-440d-b4f1-a5ca804ec0cb 1175-5326 215031 Subgenus Forcipomyia ( Saliohelea ) Wirth & Ratanaworabhan Diagnosis. Female adults: the only subgenus of Forcipomyia in the New World with fourth and fifth palpal segments completely fused, mandibular teeth absent, a hind leg tarsal ratio of 2.11–2.75 and with a single functional spermatheca; males are not diagnosable at this time.