3541 Author Judson, Sarah W. Author Nelson, C. Riley text Zootaxa 2012 2012-11-06 3541 1 118 journal article 1175­5334 505937B0-9F57-4068-82E6-8553826DD5AA Skwala DIAGNOSIS: The male hemitergal processes are knob-like ( Fig. 292 ), much shorter and stouter than those of Arcynopteryx and Megarcys . Skwala , like Diura and Filchneria , lacks an external epiproct. The female subgenital plate is trapezoidal with a slight emargination posteriorly ( Fig. 293 ). The nymph is similar to Arcynopteryx , but may be distinguished by the pleural fold extending only through abdominal segments 1 and 2 as well as the presence of serrations on both mandibles, however we did not have sufficient Mongolian specimens, particularly of the unknown A. sajanensis and S. asiatica , to confirm these purported diagnostic characteristics. DISTRIBUTION—Global: Amphi-Pacific— Regional: AOB*, IDB— Aimag: AR*, BR*,BO, SE*, OV*, UB*.