Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes
Author
Maus, Mariana Chani-Posse De
text
Zootaxa
2009
2034
31
42
journal article
10.5281/zenodo.186351
bb17e039-5346-4e01-a9ba-59b87e9e6a4c
1175-5326
186351
Philonthus quadraticeps
Boheman, 1858
(
Figs.
5, 6
in this paper; Figs.
16–21 in
Chani-Posse 2004)
Philonthus quadraticeps
Boheman,1858
: 28
;
Lynch-Arribálzaga, 1884
: 150
(characters);
Bernhauer & Schubert, 1914
: 352
(catalog);
Bruch, 1915
: 445
(catalog);
Blackwelder, 1944
: 134
(list);
Frank, 1982
; 43 (list);
Herman, 2001
: 2928
(catalog); Chani-Posse, 2004: 224–227 (characters), 231 (list).
Philonthus tenebrosus
Boheman, 1858
: 28
;
Lynch-Arribálzaga, 1884
: 149
(characters,
Argentina
);
Bernhauer & Schubert, 1914
: 357
(catalog);
Bruch, 1915
: 445
(catalog);
Scheerpeltz, 1933
: 1363
(catalog);
Blackwelder, 1944
: 135
(list);
Herman, 2001
: 2977
(catalog); Chani-Posse, 2004: 231 (list),
syn. nov.
Type
material.
Lectotype
of
Philonthus quadraticeps
(here designated;
Figs. 5
a–b), 3, “
Type
”, “Buen Ayr”, “Kinb”, “
Typus
”, “
quadraticeps
Bhn.
”, “5374 E91”, “Naturhistoriska Riksmuseet Stockholm / Loan no 834 / 06”, “
Lectotype
Philonthus quadraticeps
Boheman, 1858
/ Des. Chani-Posse de Maus 2008” (
NHRS
).
Lectotype
of
Philonthus tenebrosus
(
NHRS
) (here designated;
Figs. 6
a–b), 3, “
Type
”, “Buen Ayr”, “Kinb”, “
Typus
”, “
tenebrosus
Bhn.
”, “5373 E91”, “Naturhistoriska Riksmuseet Stockholm / Loan no 832/ 06”, “
Lectotype
Philonthus tenebrosus
Boheman, 1858
/ Des. Chani-Posse de Maus 2008” (
NHRS
).
Paralectotype
:
3, “Buen Ayr”, “Kinb”, “
tenebrosus
Bhn.
”, “5375 E91”, “Naturhistoriska Riksmuseet Stockholm / Loan no 833/06”, “
Paralectotype
Philonthus tenebrosus
Boheman, 1858
/ Des. Chani-Posse de Maus 2008” (
NHRS
).
Diagnosis.
Philonthus quadraticeps
may be identified by the following characters: head and pronotum distinctly quadrate, antennal segments 4 and 5 quadrate, eyes shorter than temples seen from above, postmandibular ridge absent, dorsal rows on pronotum each with four punctures, and transverse distance between punctures of elytra larger than diameters of punctures. For further details and illustrations see Chani-
Posse (2004)
.
Notes.
I designate the
lectotype
of
P. quadraticeps
rather than assume that the examined
type
specimen is the
holotype
since in the original description
Boheman (1858)
gave no information about the number of specimens examined. Therefore, it is possible that this name was based on more than one specimen (
ICZN 1999, Recommendation 73F
). The
lectotype
agrees with the description by
Boheman (1858)
.
In the original description of
P. tenebrosus
(
Boheman 1858
)
there is no indication about how many specimens were included in the
type
series. Both specimens listed in the
type
material agree with the description by
Boheman (1858)
. The specimen in a better condition is designated as the
lectotype
, the remaining specimen is a
paralectotype
.
Comments.
My examination of the
types
of
P. quadraticeps
and
P. tenebrosus
(
Figs. 5
a–b, 6 a–b) revealed that both are identical in all external characters and identical in characters of the male genitalia. Difference between the two nominal species as described by
Boheman (1858: 28)
results mostly from slight variation in coloration of the antennae, mouthparts and abdomen. Since the
lectotypes
of both nominal species are conspecific and these two names were simultaneously established by
Boheman (1858)
, they are considered to be subjective synonyms. Under application of the article 24.2.2 of the Code (
ICZN 1999
), I give precedence to the name
P. quadraticeps
as valid name for the species and senior subjective synonym of
P. tenebrosus
.