On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea) Author Li, Yan-Da 0000-0002-9439-202X State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China ydli@pku.edu.cn Author Tihelka, Erik 0000-0002-5048-5355 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK & wn 20250 @ bristol. ac. uk; https: // orcid. org / 0000 - 0002 - 5048 - 5355 wn20250@bristol.ac.uk Author Dahan, Loïc 0000-0001-5680-0304 O. D. Taxonomy and Phylogeny - Entomology, Royal Belgian Institute of Natural Sciences, Vautierstreet 29, B- 1000 Brussels, Belgium ydli @ pku. edu. cn; https: // orcid. org / 0000 - 0002 - 9439 - 202 X & loicdahan @ hotmail. com; https: // orcid. org / 0000 - 0001 - 5680 - 0304 loicdahan@hotmail.com Author Huang, Di-Ying 0000-0002-5637-4867 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & dyhuang @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 5637 - 4867 dyhuang@nigpas.ac.cn Author Cai, Chen-Yang 0000-0002-9283-8323 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK & cycai @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 9283 - 8323 cycai@nigpas.ac.cn text Zootaxa 2021 2021-12-17 5082 3 223 244 journal article 2910 10.11646/zootaxa.5082.3.2 3997d831-ae29-42d0-ba99-cd20cc8b7fc3 1175-5326 5788224 EFB0E4A5-6B60-4B39-A651-1C6470FD7222 Archaenosodendron explanatum Li & Cai sp. nov. ( Figs 5–8 ) Material. Holotype , NIGP177613 . The specimen seems to have been damaged before entrapment in amber, with many appendages (mouthparts, antennae, and tarsi) detached and head outstretched. Etymology. The specific name refers to its flattened body. Locality and horizon. Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District, Kachin State , Myanmar ; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian. Diagnosis. Body flattened and relatively elongate ( Fig. 7 ). Lateral regions of mentum with somewhat longitudinally arranged larger punctures ( Fig. 8B ). Anterior angles of mentum produced anteriorly ( Fig. 8B ). Antennal grooves on prosternum longer than half-length of prosternum in front of coxae ( Fig. 8D ). Outer edge of protibia with about 21–23 denticles/stout setae ( Fig. 8E–G ). Medial and lateral portions of prosternum not separated by a ridge ( Fig. 8D ). FIGURE 4. Details of Archaenosodendron cretaceum (Deng et al. , 2018) comb. nov. , holotype, CNU-COL-MA-0321, under confocal microscopy. A , Mouthparts, ventral view. B , Antenna, ventral view. C , Prothorax, ventral view, showing the incomplete boundary between the lateral medial portions of prosternum (arrowhead). D , Abdominal base, ventral view. E , Head, anterodorsal view. F , Scutellar shield, dorsal view. G–J , Fore legs. K , Mesotarsus. L , Metatibia. Abbreviations: an8–11, antennomeres 8–11; el, elytron; hd, head; lbp, labial palp; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum; v1–3, ventrites 1–3. Scale bars: 200 μm. FIGURE 5. General habitus of Archaenosodendron explanatum Li & Cai sp. nov. , holotype, NIGP177613, under incident light. A , Dorsal view. B , Ventral view. Scale bars: 1 mm. Description. Body flattened and relatively elongate, about 3.67 mm long, 1.4 mm wide. Head prognathous. Eyes not strongly protuberant. Antennal insertions widely separated and concealed from above. Antennal grooves on head well developed. Frontoclypeal suture absent. Mandibles broad, unidentate. Maxillae with setose lacinia (galea and palp detached). Mentum subtrapezoidal, widest at base; lateral regions with somewhat longitudinally arranged larger punctures; anterior angles produced anteriorly. Pronotal disc subtrapezoidal, widest at base; anterior margin with very shallow emarginations laterally, slightly protruding medially. Prosternum in front coxae transverse, not extremely shortened; medial and lateral portions of prosternum not separated by a ridge; antennal grooves on prosternum longer than half-length of prosternum in front of coxae; prosternal process relatively slender. Procoxae strongly transverse, well separated. Procoxal cavities broadly open posteriorly. Scutellar shield triangular, posteriorly acute. Elytra elongate, subparallel in the anterior half, tapering apically; anterior corners almost right. Mesoventrite short, anteriorly with well-developed mesoventral cavity and procoxal rests. Mesocoxal cavities widely separated. Metaventrite transverse, flattened; anterolateral portion with curved line delimiting the impression. Metacoxae narrowly separated. Femora with groove for receiving tibia. Tibiae expanded distally, with two apical spurs; outer edge of protibia with about 21–23 denticles/stout setae. Abdomen with five ventrites. Ventrites 1 and 2 with a pair of impressions for receiving hind legs. Anterior margin of ventrites smooth, without notches.