A new species of Halictophagus (Insecta: Strepsiptera: Halictophagidae) from Texas, and a checklist of Strepsiptera from the United States and Canada Author Kathirithamby, Jeyaraney Author Taylor, Steven J. text Zootaxa 2005 1056 1 18 journal article 10.5281/zenodo.170109 3b30e136-a327-4637-872a-82693f2775e3 1175­5326 170109 List of Strepsiptera from U.S.A. and Canada , with distribution and host records There has been no comprehensive list of Strepsiptera from USA and Canada since Kinzelbach (1971) , and one is provided here. Family Corioxenidae Kinzelbach 1970 Genus: Floridoxenos Kathirithamby & Peck 1994 : 128 Genotype: Floridoxenos monroensis Kathirithamby & Peck F. monroensis Kathirithamby & Peck 1994 : 128 ♂ Host: unknown Distribution: USA : Florida Genus Loania Kinzelbach 1970 : 99 Genotype: Loania canadensis Kinzelbach 1970 L. canadensis Kinzelbach 1970 : 99 ♂ Host: Kleidocerys Stephens ( Hemiptera : Lygaeidae ) Distribution: Canada : Ontario L. cryophila (Johnston 1976: 581) ♂ Host: unknown Distribution: USA : Georgia , Kentucky Subfamily Triozocerinae Kinzelbach 1970 : 105 Genus Triozocera Pierce 1909 Genotype: Triozocera mexicana Pierce T. mexicana Pierce 1909 : 86 ♂ Triozocera taxana Pierce 1911 : 491 Triozocera paulstana Kogan 1958 :421 Trioxocera mexicana Khalaf 1968 : 565 Triozocera mexicana Khalaf 1969 : 565 Triozocera mexicana Shepard 1979 : 217 Host: Pangaeus bilineatus (Say) ( Hemiptera : Cydnidae ) Johnson 1973, Smith & Pitts 1974 Distribution: USA : Alabama, Arizona, Florida, Kansas, Kentucky, Louisiana, Mississippi, Oklahoma, New Mexico , Texas T. vernalis Kifune and Brailovsky 1987 : 132 ♂ Host: unknown Distribution: USA : Georgia , South Carolina; Mexico Family Halictophagidae Perkins 1905 : 98 Subfamily Halictophagidae Perkins 1905 : 98 Genus Halictophagus Dale (in Curtis 1832 : 433 ) Genotype: Halictophagus curtisi Dale H. acutus Bohart 1943a : 352 ♂,Ψ, L1 Host: Draeculacephala mollipes Say , D. monerva (Ball) ( Hemiptera : Cicadellidae ) Johnston & Morrison 1979 Distribution: USA : New Mexico , Georgia , Kentucky H. americanus Perkins 1905 : 105 Ψ Agalliaphagus americanus Pierce 1908 : 80 Agallipahgus uhleri Pierce 1918 : 479 Host: Aceratagallia quadrinotata (Prov.) , A. uhleri (Van Duzee) , A. sanguinolenta (Prov.) , A. helveola ( Oman ) , A. accola Oman , A. vastitatus Oman , A. fuscohirtus Oman , A. californica (Bak.) , A. curta Oman , Eutettix tenellus (Bak.) ( Hemiptera : Cicadellidae ) Distribution: USA : Arizona, California, Colorado, Georgia , Idaho, Kansas, Maryland, Missouri, Ohio, Oregon, Utah H. barberi ( Pierce 1908: 84 ) ♂ Anthericomma barberi Pierce 1908 : 84 Host : unknown Distribution: USA : New Mexico H. bidentatus Bohart 1941 : 147 ♂ Host: Athysanella utahne Osborne , A. taxana Osborne , Deltocephalous affinis , G. & B., Eucelis frigidus , Flexamia flexulosa Ball, F. p icta Osborne, Graminella mohri Delong , Parabolocratus sp. ( Hemiptera : Cicadellidae ). Distribution: USA : Alabama, Arkansas, North Carolina, South Carolina, Georgia , Louisiana, Maryland, Mississippi, Oklahoma, Tennessee, Texas; Canada H. callosus Bohart 1943a : 350 Ψ, L1 Host: Idiocerus sp. ( Hemiptera : Cicadellidae ) Distribution: USA : Washington H. forthoodiensis Kathirithamby & Taylor , sp. n. ♂ Host: unknown Distribution: USA : Texas H. insularum ( Pierce 1908: 84 ) ♂, Ψ Dioxocera insularium Pierce 1918 : 466 Halictophagus insularium Bohart 1941 : 148 Diozocera insularum vincenti Pierce 1941 : 4 Diozocera argentinae Pierce 1941 : 5 Diozocera comstocki Pierce 1941 : 6 Diozocera comstocki elsegundinis Pierce 1941 : 10 Host: Xerophloea viridis (Fabricius) , X. vanduzeei Lawson , Dorycephalus sp. ( Hemiptera : Cicadellidae ) Distribution: USA : California, Utah H. jordani ( Pierce 1952: 5 ) ♂, Ψ, L1 Membracixenos jordani Pierce 1952 : 5 Host : Spissistylus festinus (Say) ( Hemiptera : Membracidae ) Distribution: USA : Louisiana, Texas H. mackayi ( Bohart 1937a: 103 ) ♂, Ψ, L1 Host: Aconura sp., Athysanella acuticauda (Bak,), Deltocephalous affinis G&B, Endria inimical Say. , Flexamia spp., Gramminella mohri Delong , Laevicephalaus uhleri Oman , Macrosteles fascifrons Stal, Messamia colouradensis (G&B) ( Hemiptera : Cicadellidae ) Distribution: Canada : Saskatchewan; USA : Kansas H. omani Bohart 1943a : 345 ♂, Ψ, L1 Host: Oncometopia lateralis (F.) ( Hemiptera : Cicadellidae ) Distribution: USA : Arizona, California, Kansas,Louisiana, New Mexico , Texas H. oncometopiae ( Pierce 1918: 473 ) ♂ Dacyrtocara oncometopiae Pierce 1918 : 473 Dacytrocara undata Pierce 1918 : 474 Halictophagus omcometopiae Bohart 1941 : 150 Host : Oncometopia undata (F.) ( Hemiptera : Cicadellidae ) Distribution: USA : Georgia H. philaroniae Bohart 1946 : 202 ♂, Ψ Host: Philaronia bilineata (Say) ( Hemiptera : Cercopidae ) Distribution: USA : Nebraska H. uhleri Pierce 1909 : 169 Ψ Host: Pentagramma vittifrons Uhler ( Hemiptera : Delphacidae ), Paraphlepsius irroratus (Say) (Homoptera: Cicadellidae ) Distribution: USA : Kansas, North Dakota, South Dakota. Family Elenchidae Perkins 1905 : 98 Subfamily Elenchinae Perkins 1905 : 106 Genus Elenchus Curtis 1831 : 385 Genotype: Stylops walkeri Curtis (= Elenchus tenuicornis (Kirby)) E. koebelei Pierce 1908 : 81 ♂ Elenchus tenuicornis Baumert 1959 : 400 Elenchinus heidemanni Pierce 1918 : 481 Host: Liburnia sp., Prokelisia marginata (Van Duzee), Prokelisia dolus Wilson , Sogatella kolophon (Kirkaldy) ( Hemiptera : Delphacidae ) Distribution: USA : Florida, Georgia , Kentucky, Louisiana, Maryland, Mississippi, Ohio, South Carolina Family Myrmecolacidae Saunders 1872 : 20 Genus : Caenocholax Pierce 1909 : 88 Genotype: Caenocholax fenyesi Pierce 1909 C. fenyesi texensis Kathirithamby & Johnston 2004 : S 6 ♂ Caenocholax fenyesi Pierce 1909 : 89 Caenocholax fenyesi Khalaf 1968 : 567 Caenocholax fenyesi Khalaf 1969 Caenocholax fenyesi Kathirithamby & Johnston 1992 Host: Solenopsis invicta Buren ( Hymenoptera : Formicidae ) Distribution: USA : Alabama, Arizona, California, Florida, Georgia , Louisiana, Mississippi, Texas; Mexico Family Stylopidae Kirby 1813 : 100 Subfamily Stylopinae Kirby: 1813 : 100 Genus Crawfordia Pierce 1908 : 60 Genotype: Crawfordia pulvinipes Pierce C. californica Pierce 1918 : 460 ♂ Host: Panurginus californicus Cresson ( Hymenoptera : Andrenidae ) Distribution: USA : California C. cockerelli Pierce 1909 : 155 Ψ Host: Panurginus ornatipes Cresson ( Hymenoptera : Andrenidae ) Distribution: USA : New Mexico C. labrosiformidis Pierce 1918 : 460 ♂ Host: Panurginus labrosiformidis Robertson ( Hymenoptera : Andrenidae ) Distribution: USA : Illinois

C. pulvinipes Pierce 1904: 167 ♂, Ψ, L1
Host: Pseudopanurgus innuptus (Cockerell) ,	P. labrosus Robt., P.	rudbeckiae	Robt., P. solidaginis
Robt. ( Hymenoptera : Andrenidae )
Distribution: USA: Illinois, Nebraska

Genus Halictoxenos Pierce 1908 : 82 Genotype: Halictoxenos jonesi Pierce H. crawfordi Pierce 1909 : 148 Ψ, L1 Halixenos graenicheri Pierce 1909 : 148 Halixenos zephyri Pierce 1909 : 148 Halixenos versati Pierce 1909 : 148 Halixenos sparsi Pierce 1909 : 148 Host: Halictus bruneri Crawford , H. albipennis Robt. , H. versatus Robt. , H. zephyrus Robt. , H. incompletes Crawford , H. punctaroventris Crawford aff. H. nevadensis Cockerell ( Hymenoptera : Halictidae ) Distribution: USA : Alabama, California, Illinois, Nebraska, North Carolina, Oklahoma, Wisconsin H. jonesi Pierce 1908 ♂ Host: Dialictus imitatus ( Hymenoptera : Halictidae ) Distribution: USA : Louisiana, Texas H. nymphaeri Pierce 1911 : 496 Ψ Host: Halictus nymphaerum Robt. ( Hymenoptera : Halictidae ) Distribution: USA : Illinois H. schwarzi ( Pierce 1908: 83 ) ♂ Apractelytra schwarzi Pierce 1909 : 83 Host: Halictus geminus Sandhouse ( Hymenoptera : Halictidae ) Distribution: USA : District of Columbia H. viridulae Pierce 1911 : 496 Ψ Host: Augochlora viridula Smith ( Hymenoptera : Halictidae ) Distribution: USA : Illinois Genus Hylecthrus Saunders 1850 : 57 Genotype: Hylecthrus rubi Saunders H. californicus Bohart 1941 : 140 ♂, Ψ, L1 Host: Hylaeus sp. ( Hymenoptera : Colletidae ) Distribution: USA : Arizona, California Colorado, Genus Eurostylops Bohart 1943a : 12 Pseudostylops Bohart 1943b : 134 Genotype: Eurostylops desertorum Bohart E. desertorum ( Bohart 1941: 134 ) Ψ, L1 Host: Halictoides boregoensis Michener ( Hymenoptera : Halictidae ) Distribution: USA : California E. conanthalicti Kinzelbach 1971 : 165 Ψ, L1 Host: Conanthalictus retito Ger., Conathalictus sp. ( Hymenoptera : Halictidae ) Distribution: USA : California E. sierrensis Bohart 1978 : 101 Ψ Host: Dufourea trochantera Bohart ( Hymenoptera : Halictidae ) Distribution: USA : California E. tetonensis Bohart 1943b : 13 Ψ Host: Halictodes maurus Cresson ( Hymenoptera : Halictidae ) Distribution: USA : Wyoming Genus Stylops Kirby 1802 : 112 Genotype: Stylops melitte Kirby S. apicalis Bohart 1937a : 54 Ψ, L1 Host: Andrean saccata Viereck ( Hymenoptera : Andrenidae ) Distribution: USA : California, Oregon S. bipunctatae Pierce 1909 : 98 Ψ Host: Andrena bipunctata Cresson ( Hymenoptera : Andrenidae ) Distribution: USA : Indiana, Nebraska, Wisconsin S. bisalicidis Pierce 1918 : 446 Ψ Host: Andrena bisalicis Viereck ( Hymenoptera : Andrenidae ) Distribution: USA : Alabama, North Dakota S. bruneri Pierce 1909 : 98 Ψ Stylops hartfordensis Pierce 1909 : 103 Stylops nasoni Pierce 1909 : 104 Stylops andrenoides Pierce 1911 : 493 Stylops salictariae Pierce 1918 : 449 Host: Andrean salictaria Robt. , A. ililnoiensis Robt. , A. nasoni Robt. , A. andrenoides Cresson ( Hymenoptera : Andrenidae ) Distribution: USA : Alabama, Indiana, Nebraska, Wisconsin S. californicus Pierce 1909 : 99 Ψ Host: Andrena subtilis Smith ( Hymenoptera : Colletidae ) Distribution: USA : California S. claytoniae Pierce 1909 : 99 Ψ Stylops imitatrix Pierce 1909 : 104 Stylops vierecki Pierce 1909 : 110 Host: Andrena imitatrix Cresson , A. claytonae Robt. ( Hymenoptera : Andrenidae ) Distribution: USA : Alabama, Georgia , Illinois, Oklahoma, Texas S. cornii Pierce 1909 : 100 Ψ, L1 Stylops graenicheri Pierce 1909 : 103 Host: Andrena commoda Smith , A. nivailis Smith ( Hymenoptera : Andrenidae ) Distribution: USA : Wisconsin S. crawfordi Pierce 1909 : 100 ♂, Ψ Host: Andrena crawfordi Viereck ( Hymenoptera : Andrenidae ) Distribution: USA : Texas S. cressoni Pierce 1909 : 102 Ψ, L1 Host: Andrena cressoni Robt. ( Hymenoptera : Andrenidae ) Distribution: USA : Maine S. cuneiformis Bohart 1936 : 16 ♂ Host: Andrena ( Hymenoptera : Andrenidae ) Distribution: USA : California S. elongata Bohart 1937a : 53 Ψ, L1 Host: Andrena oenothera Timberlake , A. blaisdelli Cockerell Distribution: USA : California S. heterocingulata Bohart 1937a : 55 Ψ, L1 Host: Andrena pensilis Timberlake , A. angustitarsata huardi Viereck , A. opaciventris Cockerell ( Hymenoptera : Andrenidae ) Distribution: USA : California, Oregon S. hippotes Pierce 1909 : 103 Ψ Host: Andrena hippotes Robt. ( Hymenoptera : Andrenidae ) Distribution: USA : Ohio S. medionitans Pierce 1918 : 450 Ψ Host: Andrena medionitans, Cockerell , A. scurra Viereck , Andrena sp. ( Hymenoptera : Andrenidae ) Distribution: USA : California, Oregon, Colorado S. moestae Pierce 1918 : 443 Ψ Host: Andrena moesta Smith ( Hymenoptera : Andrenidae ) Distribution: USA : Washington S. multiplicatae Pierce 1909 : 104 Ψ Host: Andrena multiplicata Cockerell ( Hymenoptera : Andrenidae ) Distribution: USA : Wisconsin S. neonanae Pierce 1918 : 454 Ψ Host: Andrena nernana Vierbeck ( Hymenoptera : Andrenidae ) Distribution: USA : Georgia S. nubeculae Pierce 1909 : 105 Ψ Host: Andrena nubecula Smith ( Hymenoptera : Andrenidae ) Distribution: USA : Colorado S. nudae Pierce 1911 : 495 Ψ Host: Andrena nuda Robt. ( Hymenoptera : Andrenidae ) Distribution: USA : Illinois S. oklahomae Pierce 1909 : 110 Ψ Host: Andrena flavoclypeata miserabilis Cresson ( Hymenoptera : Andrenidae ) Distribution: USA : Oklahoma S. packardi Pierce 1909 : 105 ♂ Host: Andrena placida Smith ( Hymenoptera : Andrenidae ) Distribution: USA : Massachusetts S. pacificus Bohart 1936 : 15 ♂, Ψ, L1 Host: Andrena complexa Viereck , A. suavis Timberlake , A. cuneilabris Viereck , Panurginus melanocephalus (Cochrell) ( Hymenoptera : Andrenidae ) Distribution: USA : California, Oregon S. salicifloris Pierce 1909 : 106 Ψ Stylops centroclarus Bohart 1937a : 50 Host: Andrena salicifloris Cockerell , A. quintilormis Viereck ( Hymenoptera : Andrenidae ) Distribution: USA : California, Washington S. shannoni Pierce 1918 : 457 ♂ Stylops shannoni Kenner 2002 : 160 ♂, Ψ, L1 Host: Andrena hippotes Robertson ( Hymenoptera : Andrenidae ) Distribution: USA : Maryland; Canada S. solidulae Pierce 1909 : 107 ♂, Ψ Host: Andrena solidula Viereck ( Hymenoptera : Andrenidae ) Distribution: USA : Washington S. sparsipilosae Pierce 1909 : 108 Ψ Host: Andrena sparsipilosa Viereck ( Hymenoptera : Andrenidae ) Distribution: USA : Maine S. subcandidae Pierce 1909 : 108 Ψ, L1 Host: Andrena subcanidida ( Hymenoptera : Andrenidae ) Distribution: USA : California S. swenki Pierce 1909 : 108 Ψ, L1 Stylops asteridis Pierce 1911 : 494 Host: Andrena asteris Robertson, A. aff. lincolnella Cockerell ( Hymenoptera : Andrenidae ) Distribution: USA : Illinois, Nebraska, Pennsylvania S. timberlakei Bohart 1936 : 14 ♂, Ψ Host: Andrena macrocephala Cockerell ( Hymenoptera : Andrenidae ) Distribution: USA : California S. vandykei Bohart 1936 : 11 ♂, Ψ, L1 Host: Andrena perimelas Cockerell , A. carliniformis Cockerell ( Hymenoptera : Andrenidae ) Distribution: USA : California, Oregon Subfamily Xeninae Saunders 1872 : 20 Genus Pseudoxenos Saunders 1872 : 44 Gentotype: Pseudoxenos schaumi Saunders P. arvensidis Pierce 1911 : 499 Ψ Host: Odynerus ( Rygchium) arvensis Saussure , O. ( Rygchium ) sulphurous Saussure ( Hymenoptera : Vespidae ) Distribution: USA : Illinois, California P. bishoppi ( Pierce 1909: 139 ) Ψ, L1 Monobiaphila bishoppi Pierce 1909 : 139 Host : Monobia quadridens (L) ( Hymenoptera : Vespidae ) Distribution: USA : Texas P. erynnidis Pierce 1911 : 499 Ψ Host: Pachodynerus erynnys (Lepeletier) ( Hymenoptera : Vespidae ) Distribution: USA : Florida P. foraminati Pierce 1911 : 499 Ψ Host: Rygchium foraminatum (Saussure) ( Hymenoptera : Vespidae ) Distribution: USA : New Jersey P. hookeri ( Pierce 1909: 139 ) Ψ, L1 Leionotoxenos hookeri Pierce 1909 : 139 Host: Rygchium annulatum annulatum (Say) ( Hymenoptera : Vespidae ) Distribution: USA : California, Illinois, Texas P. j o n e s i ( Pierce 1909: 138 ) Ψ Leionotoxenos joensi Pierce 1909 : 138 Host: Stenodynerus fulvipes (Saussure) ( Hymenoptera : Vespidae ) Distribution: USA : Lousiana, North Carolina, Texas, P. louisianae ( Pierce 1909: 138 ) Ψ, L1 Leinontoxenos louisianae Pierce 1909 : 138 Pseudoxenos histrionis Pierce 1911 : 500 Pseudoxenos pedestridis Pierce 1911 : 500 Host: Stenodynerus histrio (Lepeletier) , S. pedestris (Sassure) , S. vagus (Sassure) ( Hymenoptera : Vespidae ) Distribution: USA : Nebraska P. neomexicanaus Pierce 1918 : 463 ♂ Host: Stenodynerus taos (Cresson) ( Hymenoptera : Vespidae ) Distribution: USA : New Mexico P. ro b e r s o n i Pierce 1911 : 501 Ψ Host: Stenodynerus annonia histrionalis (Robertson) ( Hymenoptera : Vespidae ) Distribution: USA : Illinois P. tigridis Pierce 1911 : 501 Ψ Host: Odynerus tigris Saussure ( Hymenoptera : Vespidae ) Distribution: USA : Illinois Genus Xenos Rossi 1793 : 49 Genotype: Xenos vesparum Rossius X. hunteri ( Pierce 1909: 130 ) ♂, Ψ Acroschismus hunteri Pierce 1911a : 130 Host : Polistes sp. ( Hymenoptera : Vespidae ) Distribution: USA : Texas X. kifunei Cook & Mathison 1997 : 256 ♂ Host: Polistes comanchus navajoe ( Hymenoptera : Vespidae ) Distribution: USA : Arizona X. pallidus Brues 1903: 246 ♂, Ψ Acroschismus hubbardi Pierce 1908 : 84 Acroschismus pallidus texensis Pierce 1909 : 127 Host: Polistes sp., Polistes canadensis annularis (Linneaus) ( Hymenoptera : Vespidae ) Distribution: USA : Alabama, Nebraska, South Carolina, Texas X. pecki Kirby 1813 : 100 ♂ Xenos wheeleri Pierce 1908 : 80 Xenos bruesi Pierce 1909 : 124 Acroschismus pecosensis Pierce 1909 : 128 Acroschismus bowditchi Pierce 1909 : 130 Acroschismus texani Pierce 1909 : 132 Acroschismus maximus Pierce 1909 : 132 Xenos auriferi Pierce 1911 : 498 Xenos californicus Pierce 1918 : 489 Host: Polistes fuscatus (F.), Polistes metricus Say ( Hymenoptera : Vespidae ) Distribution: USA : Alabama, Georgia , Hawaii, South Carolina X. rubiginosi ( Pierce 1909: 132 ) Ψ Acroschismus rubiginosi Pierce 1909 : 132 Host : Polistes rubiginosus Lepeletier ( Hymenoptera : Vespidae ) Distribution: USA : Lousiana Subfamily Paraxeninae Kinzelbach 1971 : 162 Genus Paraxenos Saunders 1872 : 45 Genotype: Paraxenos erberi Saunders P. auripedis ( Pierce 1911: 503 ) Ψ Ophthalmochlus auripedis Pierce 1911 : 503 Host : Isodontia auripes (Fernald) ( Hymenoptera : Sphecidae ) Distribution: USA : Pennsylvania, Maryland P. duryi ( Pierce 1908: 83 ) ♂ Ophthalmochlus duryi Pierce 1908 : 83 Host : Prionyx atratus (Lepeletier) ( Hymenoptera : Sphecidae ) Distribution: USA : Ohio, Louisiana P. luctusae ( Pierce 1911: 502 ) Ψ Eupathocera luctusae Pierce 1911 : 502 Host: Podalonia argentiforns (Cresson) , P. violaceipennis (Lepeletier) , P.v. luctusae (Smith) ( Hymenoptera : Sphecidae ) Distribution: USA : Maryland P. lugubris ( Pierce 1908: 83 ) ♂ Eupathocera lugubris Pierce 1908 : 83 Eupathocera pruinosae Pierce 1909 : 143 Eupathocera pictipennidis Pierce 1911 : 502 Eupathocera vulgardis Pierce 1911 : 503 Host: Sphex sp. ( Hymenoptera : Sphecidae ) Distribution: USA : Colorado, Illinois, Ohio P. westwoodi ( Templeton 1841: 51 ) ♂ Xenos westwoodi Templeton 1841 Xenos smithii Heyden 1867 : 398 Paraxenos westwoodi Saunders 1872 Hamilops westwoodi Pierce 1908 Hamilops bishoppi Pierce 1909 : 146 Hamilops ashmeadi Pierce 1909 : 146 Host: Sphex ichneumoneus (Linneaus) ( Hymenoptera : Sphecidae ) Distribution: USA : California, Gerogia, Illinois, North Carolina, Ohio, Texas