A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia Author Zonstein, Sergei L. text European Journal of Taxonomy 2024 2024-10-24 967 1 185 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459 journal article 10.5852/ejt.2024.967.2699 2118-9773 13990819 C08B8027-50CC-417E-BCD4-5183B9FF6738 Raveniola ignobilis sp. nov. urn:lsid:zoobank.org:act: 17DD6055-088F-4BEF-A1D6-DD4686AF191C Figs 9 , 43 , 91 , 118 , 145 , 175 , 198 , 210 , 237 , 267 , 297 , 315 , 358–359 , 403–408 , 509 , 573–574, 629– 631, 752 Diagnosis Due to its relatively short copulatory bulb, males of Raveniola ignobilis sp. nov. resemble the holotype of R . dolosa sp. nov. , but differ from the latter in possessing two raised opposite keels in the proximal part of the embolus ( Figs 403–408 cf. Figs 397–399 ). The conspecific females are distinguishable by a specific structure of their spermathecae, with long inner and outer branches set very close to each other ( Fig. 509 ) vs differently arranged branches in other related species, where these branches are either shorter ( Figs 500–503 , 524–525 ) or broader spaced ( Figs 495–499 , 520–523 ). Etymology The specific epithet is a Latin adjective meaning ‘obscure’, ‘inglorious’ and referring to a rather middling (or averaged) appearance of these congeners. Material examined Holotype TAJIKISTAN • ♂ ; Hazratisho Mts , Kapar (Sangdara) Gorge , environs of Sangdara Village ; 38°22′ N , 70°10′ E ; 1650 m a.s.l. ; 15 Oct. 1987 ; S. Zonstein leg.; SMNH . Paratypes ( 4 ♂♂ , 1 ♀ , 1 ♀ subad.) TAJIKISTAN • 4 ♂♂ , 1 ♀ ; same collection data as for preceding; 1450–1800 m a.s.l. ; SMNH • 1 ♀ subad.; same collection data as for preceding, Iokunj Gorge ; 38°23′ N , 70°09′ E ; 1700 m a.s.l. ; 18 May 2002 ; S. Zonstein leg.; SMNH . Additional material ( 1 ♀ subad., 1 juv. ) TAJIKISTAN • 1 ♀ subad.; Hazratisho Mts. , 25 km E of Muminabad Town ; 24 May 1966 ; E.M. Andreeva leg.; MIZW • 1 juv. ; Darvaz Mts , Hirakdara Canyon between Kalai-Husain and Safedoron Villages; 2300 m a.s.l.; Juniperus park forest; 4 Jul. 1970 ; E.M. Andreeva leg.; MIZW . Description Male ( holotype ) HABITUS . See Fig. 9. MEASUREMENTS . TBL 14.40, CL 5.96, CW 5.59, LL 0.45, LW 0.89, SL 3.16, SW 2.87. COLOUR . Carapace and leg I from femur to basal metatarsus intensely brownish orange; other parts of leg I, and entire palps and legs II–IV, as well as sternum, labium and maxillae lighter brownish orange; eye tubercle with eyes surrounded with partially fused blackish rings, chelicerae light cherry red, abdomen pale greyish brown, dorsally with diffuse and weakly developed brownish chevron-like pattern; book-lungs and spinnerets very pale greyish yellow. CEPHALOTHORAX . Carapace and chelicerae as shown in Fig. 91 . Clypeus and eye group as in Fig. 145 . Eye diameters and interdistances: AME 0.15(0.20), ALE 0.27, PLE 0.22, PME 0.16; AME–AME 0.14(0.09), ALE–AME 0.09(0.06), ALE–PLE 0.09, PLE–PME 0.04, PME–PME 0.32. Anterior cheliceral edge with unmodified setae; rastellum not developed. Each cheliceral furrow with 9 promarginal teeth and 5–6 mesobasal denticles. Presumed intercheliceral tumescence poorly discernible, small, maculate and setose ( Fig. 198 ). Sternum, labium and maxillae as shown in Fig. 210 . Maxillae with ca 60 cuspules each. LEGS . Tibia and metatarsus I as in Figs 267 , 297 . Scopula: entire and distal on metatarsi I–II; entire on tarsi I–II; widely divided on tarsi III–IV. Trichobothria: 2 rows of 9–10 each on tibiae, 13–15 on metatarsi, 12–13 on tarsi, 10 on cymbium. PTC I–II and III–IV with 8–9 and 10–12 teeth on each margin, respectively. SPINATION . Palp: femur d4, pd1, rd1; patella pd1; tibia d4, p3, r1, v6; cymbium d3(5)+20–25 spikes. Leg I: femur d2+2 bristles, pd3; patella p1; tibia p2, pv2, rv2+ 2M. Leg II: femur d2+2 bristles, pd3; patella p1; tibia p3, v7; metatarsus p1, v5. Leg III: femur d4, pd3, rd2; patella p2(1), r1; tibia d4, p4, r3, v7; metatarsus d4, p4, r3, v6. Leg IV: femur d3, pd3, rd2; patella p2, r1; tibia d3, p4, r3, v7; metatarsus d1, p4, r3, v8. Tarsi I–IV aspinose. PALP . Tibia, cymbium and copulatory bulb as shown in Figs 358–359 . Embolus with long basal portion provided with low keel and short hooked apical part ( Figs 403–405 ). SPINNERETS . See Fig. 573. PMS: length 0.48, diameter 0.14. PLS: maximal diameter 0.51; length of basal, medial and apical segments 0.80, 0.49, 0.37; total length 1.66; apical segment triangular. LEG MEASUREMENTS . ♂(♀)
Femur Patella Tibia Metatarsus Tarsus Total
Palp 3.74 (3.74) 1.89 (2.02) 2.92 (2.42) – 0.97 (2.11) 9.52 (10.29)
Leg I 6.09 (4.84) 3.13 (2.96) 4.80 (3.52) 4.65 (2.64) 2.59 (1.77) 21.26 (15.73)
Leg II 5.70 (4.41) 2.81 (2.59) 4.58 (3.18) 4.39 (2.72) 2.55 (1.78) 20.03 (14.68)
Leg III 4.97 (3.72) 2.09 (2.01) 3.53 (2.45) 4.36 (3.17) 2.57 (1.93) 17.52 (13.28)
Leg IV 6.03 (4.87) 2.49 (2.62) 4.79 (3.51) 6.52 (4.55) 3.01 (2.43) 22.84 (17.98)
Female ( paratype ) HABITUS . See Fig. 43. MEASUREMENTS . TBL 16.60, CL 6.14, CW 5.58, LL 0.69, LW 1.19, SL 3.17, SW 2.86. COLOUR . Similar to that of male, but dorsal abdominal chevron-like pattern darker and better developed. CEPHALOTHORAX . Carapace and chelicerae as shown in Fig. 118 . Clypeus and eye group as in Fig. 175 . Eye diameters and interdistances:AME 0.16(0.22), ALE 0.30, PLE 0.17, PME 0.16; AME–AME 0.11(0.05), ALE–AME 0.10(0.07), ALE–PLE 0.09, PLE–PME 0.03, PME–PME 0.43. Cheliceral rastellum absent. Each cheliceral furrow with 8–9 promarginal teeth and 5–6 mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 237 . Maxillae with 56–57 cuspules each. LEGS . Tibia and metatarsus IV as shown in Fig. 315 . Scopula: distal on metatarsi I–II; entire on palpal tarsus and tarsi I–II; sparse and widely divided by setae on tarsus III; rudimentary on tarsus IV. Trichobothria: 2 rows of 7–8 each on tibiae, 13–15 on metatarsi, 12–14 on tarsi. Palpal claw with 4 promarginal teeth. PTC I–II and III–IV with 4–5/5–6 and 6–7/8–9 teeth on inner/outer margins, respectively. SPINATION . All femora with one basodorsal spine and 3–4 median and/or apical bristles; palpal patella, patella I, and tarsi I–IV aspinose. Palp: femur pd1; tibia v7(6); tarsus v3(2). Leg I: femur pd2; tibia v3; metatarsus v6. Leg II: femur pd3; patella p1; tibia p2, v3; metatarsus v6. Leg III: femur pd3, rd3(2); patella p2, r1; tibia d1, p2, r2, v7; metatarsus d4, p3, r3, v7. Leg IV: femur pd1, rd1; patella p1, r1; tibia d1, p3, r3, v7; metatarsus d1, p4, r4, v8. SPERMATHECAE . Each of paired spermathecae V-shaped with relatively short and wide base carrying two closely arosing, equally thick, long and weakly diverging branches ( Fig. 509 ). SPINNERETS . See Fig. 574. PMS: length 0.61, diameter 0.23. PLS: maximal diameter 0.63; length of basal, medial and apical segments 1.06, 0.56, 0.28; total length 1.90; apical segment triangular. Variation Carapace length in the paratype males (n =4) varies from 5.49 to 6.96. Some variations in the structure of the copulatory bulb as shown in Figs 406–408 . Ecology The spiders were collected from their retreats under stones in different montane biotopes, mostly from the mixed woodlands at 1400–2300 m , dominated by Juglans regia and Juniperus seravschanica ( Figs 629– 631 ). Distribution Tajikistan : Hazratisho and Darvaz Mts. See Fig. 752 .