Identity of species-group taxa of the Western Palaearctic Clytrini (Coleoptera: Chrysomelidae) described by Maurice Pic and Louis Kocher
Author
Bezdċk, Jan
Author
Regalin, Renato
text
Acta Entomologica Musei Nationalis Pragae
2015
suppl.
2015-10-15
55
1
114
journal article
9543
10.5281/zenodo.4272771
77c874cd-4f85-4746-8466-3ca09e2c2b8d
0374-1036
4272771
urn:lsid:zoobank.org:pub:A431893C-11D8-4480-ABE1-BB65A823C734
Tituboea biguttata
(
Olivier, 1791
)
(
Figs 68–70
)
Clytra biguttata
Olivier, 1791: 34
(original description).
Antipa
biguttata
var.
vagenotata
Pic, 1939: 18
(original description).
Antipa
bicoloripes
Pic, 1942a: 78
(original description) [published on
31
st
May 1942
according to Ballerio, pers. comm. 2015],
syn. nov.
Antipa bicoloripes
Pic, 1942b:10
(duplicate description, junior primary homonym and objective synonym) [published on
18
th
Aug 1942
according to the title page of the publication],
syn. nov.
Antipa
tredecimpunctata
var.
ssavipennis
Pic, 1950: 16
(original description).
Antipa
biguttata
var.
legionis
Kocher, 1959: 12
(unavailable infrasubspeciħc name).
Type
localities.
Clytra biguttata
: ‘Espagne’. Antipa
biguttata
var.
vagenotata
: ‘Tunis’.
Antipa
bicoloripes
: ‘Adrar des Iforas’ [northeastern
Mali
]. Antipa
tredecimpunctata
var.
ssavipennis
: ‘
Maroc
: Grand-Atlas’. Antipa
biguttata
var.
legionis
: ‘Ziz, près de Rich; près de Bou-Arfa’ [=
Morocco
, Ziz river, near Er-Rich; near Bouarfa].
Type
material examined.
Clytra biguttata
: not examined.
Antipa
biguttata
var.
vagenotata
: not examined. Not found in
MNHN
.
Antipa
bicoloripes
:
SYNTYPES
: 1♂, ‘
Adrar
des
Iforas
/
Od Tin Biden
/ (s. sahélo-sahar.) /
6-8 aout 1941
/
Volkonsky
[w, h] //
Antipa
(1) [w, h] // désiré [w, h] //
Antipa
/ bicoloripes / n sp [w, h] //
TYPE
[r, p]’ (
MNHN
– coll.
Pic
)
;
1 ♀
, ‘
Kidal
/
Adrar
des
Iforas
/
aout 1941
/
Volkonsky
[w, h] //
TYPE
[r, p]’ (
MNHN
– coll.
Pic
)
;
1 ♂, ‘
Kidal
/
Adrar
des
Iforas
/
aout 1941
/
Volkonsky
[w, h] //
Antipa
/
bicoloripes Pic
/ c.types [w, h] // décrit in / l
̓
Echange No
489 / 1942, p. 10 [w, h]’ (
MNHN
– coll.
Peyerimhoff
)
;
1 ♀
, ‘
Adrar
des
Iforas
/
Od Tin Biden
/ (s. sahélo-sahar.) /
6-8 aout 1941
/
Volkonsky
[w, h]’ (
MNHN
– coll.
Peyerimhoff
)
;
1 ♀
, ‘
Kidal
/
Adrar
des
Iforas
/
aout 1941
/
Volkonsky
[w, h]’ (
MNHN
– coll.
Peyerimhoff
)
.
Antipa
tredecimpunctata
var.
ssavipennis
: not examined.
Original material of infrasubspeciħc entity.
Antipa
biguttata
var.
legionis
: 1♂, ‘Rich (Ziz) [h]/ SUD
MAROC
[p] / (Roulleau) 5.53 [w, h] // [small blank round blue label] //
var. legionis
/ m. [h] /
Kocher
det. [w, p] //
HOLOTYPE
[r, p]’ (
PJCP
);
1 ♀
,‘foud ouet [h] / Bou-Arfa / Mar. Ortal [w, p] // Maroc[p] 30-5-54 [h] / A. REYMOND [w, p] //
♀
[w, p] // [small blank round blue label] //
PARATYPE
[r, p]’ (
PJCP
).
Distribution.
Mediterranean species (for details see
REGALIN & MEDVEDEV 2010b
).
Comments.
Tituboea biguttata
is one of the most common and widespread Mediterranean clytrines with extensive variability of dorsal pattern. This species evidently forms many local populations which can slightly differ also in the shape of the aedeagus and male protarsi.
The
type
material of
T. biguttata
was not traced in MNHN and its deposition is unknown. However, there is no doubt about the identity of the species based on the description (
OLIVIER 1791
) and colour picture in
OLIVIER (1808)
.
The
type
material of
var.
vagenotata
was not found in MNHN and the deposition is unknown to us. We left this taxon in synonymy with
T. biguttata
.
According to
KOCHER (1959: 9
, note 2),
A. tredecimpunctata
var.
ssavipennis
was described based on one female (
holotype
) deposited in Institut scientiħque chériħen in
Rabat
. In PJCJ only non-type specimen mentioned by
KOCHER (1959: 12)
was found. However, there is no doubt about its synonymy with
Tituboea biguttata
.
Antipa
biguttata
var.
legionis
was described in a publication containing also descriptions of subspecies (
KOCHER 1959
), thus it became infrasubspeciħc according to Article 45.6.4 of the Code (
ICZN 1999
). Two original specimens are deposited in PJCP and bear ‘holotype’ and ‘paratype’ labels, but these were added later by Jolivet (Jolivet, pers. comm. 2014). This variety is conspeciħc with
T. biguttata
.
The type series of
A. bicoloripes
is split into two parts: two
syntypes
are placed in Pic
̓
s collection and additional three
syntypes
are in Peyerimhoff
̓
s collection. The colouration of the specimens, particularly on pronotum, resembles that of
T. tredecimpunctata
, however examination of aedeagus conħrmed that
A. bicoloripes
is conspeciħc with
T. biguttata
. Protarsi I in the male
syntypes
of
A. bicoloripes
are slightly shorter than in typical
T. biguttata
, however we think that such difference can occur in local marginal population of this widely distributed species.
Antipa
bicoloripes
was described twice based on the same type specimens (
PIC 1942a
,b), thus, both names are objective synonyms and are also proposed as new synonyms of
T. biguttata
.