New species and new records of stomatopod Crustacea from the Philippines Author Ahyong, Shane T. text Zootaxa 2004 793 1 28 journal article 10.5281/zenodo.169642 81f758e2-353b-442e-85c1-104b675ea597 1175­5326 169642 DB1B08BE-30CC-419A-A5DD-583A072449D7 Carinosquilla balicasag sp. nov. ( Figure 3 ) Material examined. HOLOTYPE : AM P67177, female (TL 82 mm ), Balicasag Island, P. Ng et al. from local fishermen, 26 Jul 2003 . PARATYPES : ZRC 2003.0308, 1 male (fragmented, CL 24.5 mm ), type locality, P. Ng et al. from local fishermen, 25–30 Jul 2003 ; ZRC 2003.0307, 1 male (fragmented, CL 24.8 mm ), type locality, P. Ng et al. from local fishermen, 25–30 Jul 2003 ; ZRC 2004.0615, 1 male (TL 106 mm ), type locality, P. Ng et al. from local fishermen, 2 Mar 2004 . Diagnosis. Eyestalk with short carinae. Ocular scales undivided. Anterior bifurcation of median carina of carapace opening anterior to dorsal pit. Raptorial claw dactylus with 5 teeth; outer surface of merus with 2 longitudinal carinae. Mandible with palp. TS5 supplementary carinae longitudinal or oblique, not transverse. TS6–8 and AS 1–5 with most or all supplementary dorsal carinae posteriorly spined above level of intermediate carinae. AS 1– 5 with 1 or 2 (usually 2) supplementary carinae armed between intermediate and lateral carinae; supplementary carinae between lateral and marginal carinae unarmed. AS 6 submedian carinae tricarinate; supplementary carinae between submedian and intermediate carinae unarmed. Telson prelateral lobe with blunt apex; with 1 lateral denticle. Uropodal exopod distal segment dark on proximal half or one­third. Description. Cornea strongly bilobed; carinae on eyestalk short, irregular; CI 412– 448. Ocular scales entire, apices truncate, not bifurcate. A1 peduncle length 1.04–1.13 CL. A1 somite dorsal processes with acute triangular apices; directed anterolaterally. A2 scale length 0.70–0.75 CL; with irregular carinae proximally. Rostral plate as long as broad; lateral margins convergent, upturned, sinuous; apex rounded; median carina flanked by long supplementary carinae. Carapace anterior width 0.42–0.44 CL; anterior bifurcation of median carina opening anterior to dorsal pit; dorsum covered with numerous closely spaced longitudinal carinae; anterolateral spines short, not reaching anteriorly to level of base of rostral plate. Raptorial claw dactylus with 5 teeth; outer surface of propodus smooth; carpus with undivided dorsal carina; outer surface of merus with 2 longitudinal carinae, lower carina flanked ventrally by distinct groove; ischium unarmed; basis with short ventrally directed spine at articulation with ischium. Mandibular palp 3­segmented. MXP1–4 each with epipod. TS5 lateral process bilobed; anterior lobe a short spine directed anterolaterally; posterior lobe with rounded apex, directed laterally. TS6 lateral process bilobed; anterior lobe broad, quadrate, apex truncate; posterior lobe broad, triangular, anterior margin carinate, sinuous, apex blunt. TS7 lateral process bilobed; anterior lobe triangular, blunt; posterior lobe larger than anterior lobe, broad, triangular, anterior margin convex, carinate, apex angular. TS8 anterolateral margin triangular, apex angular; sternal keel angular. FIGURE 3. Carinosquilla balicasag sp. nov. A–K, holotype female TL 82 mm (AM P67177). L, paratype male, TL 106 mm (ZRC 2004.0615). A, anterior. B, dorsal process of A1 somite, right lateral. C, carapace, posteromedian portion. D, right raptorial claw, lateral view. E, TS5–7 dorsum and right lateral processes. F, TS5, right lateral view. G, TS8 sternal keel, right lateral view. H, AS1, right lateral view. I, AS5–6, telson and uropod. J, AS6 and telson, ventral. K, right uropod, ventral. L, pleopod 1 endopod. Scale A–K = 5.0 mm, L = 2.5 mm. TS5 with numerous dorsal carinae, longitudinal or oblique, not transverse except for carinae along anterior margin; most carinae posteriorly spined. TS6–8 and AS 1–5 with most or all supplementary dorsal carinae posteriorly spined above level of intermediate carinae. AS 1–5 with 1 or 2 (usually 2) supplementary carinae armed between intermediate and lateral carinae; supplementary carinae between lateral and marginal carinae unarmed. Articular membrane between AS 5 and 6 lined with small spines. AS 6 submedian carina tricarinate; posterior margin with 2 short spinules lateral to submedian spine; supplementary carinae between submedian and intermediate carinae unarmed; sternum with anterior and posterior transverse carina, median carina flanked by 4–5 sinuous carinae, some uniting laterally. Abdominal somites with normal complement of carinae spined as follows: submedian 1–6, intermediate 1–6, lateral 1–6, marginal 1–5. Telson about as long as broad; prelateral lobe longer than margin of lateral tooth, apex blunt; dorsolateral surface with numerous supplementary longitudinal carinae. Telson denticles: submedian 4–5, intermediate 9–11, lateral 1; ventral surface with long postanal carina, numerous longitudinal supplementary carinae. Uropodal protopod densely carinate; outer margin smooth; inner margin with 9–13 slender spines; with short ventral spine anterior to endopod articulation. Protopod terminal spines with lobe on outer margin of inner spine rounded, narrower than adjacent spine, proximal margin straight or slightly concave. Exopod proximal segment dorsally and ventrally carinate; outer margin with 10 or 11 movable spines, distalmost not exceeding midlength of distal segment; distoventral margin with 2 spines, outer longest. Exopod distal segment shorter than proximal segment; dorsally and ventrally carinate; dark on proximal half or one­third; dorsally and ventrally carinate. Endopod dorsally and ventrally carinate. Colour in alcohol. Overall pale tan with dark­brown grooves and carinae. AS 2 and AS 5 with pair of dark­brown submedian patches. Telson with dark band lateral margins extending from prelateral lobe to inner intermediate denticles. Terminal spines of uropodal protopod and distal third of uropodal endopod dark brown. Proximal segment of uropodal exopod with dark brown patch extending onto proximal third of distal segment. Distal two­thirds of uropodal exopod pale. Etymology. Named after the type locality, Balicasag Island; used as a noun in apposition. Measurements of holotype . TL 82 mm , CL 19.70 mm , anterior carapace width 8.60 mm , cornea width 4.40 mm , A1 peduncle 22.30 mm , A2 scale 13.90 mm . Remarks. Carinosquilla balicasag sp. nov. resembles C. australiensis Ahyong, 2001 from eastern Australia in almost all respects but differs in having none of the supplementary carinae between the submedian and intermediate carinae of AS 6 armed posteriorly, and in having the distal segment of the uropodal exopod darkly pigmented only on the proximal third instead of its entirety. As in C. australiensis , one or more spinules may be present along the posterior margin of AS 6 lateral to the submedian carinae. Distribution. Known only from Balicasag Island.