The morphology of the pupae of six species of Philonthina (Coleoptera, Staphylinidae, Staphylinini) with taxonomic remarks Author Pietrykowska-Tudruj, Ewa Author Staniec, Bernard text Zootaxa 2011 2865 53 67 journal article 10.5281/zenodo.202575 e6fc0416-99c9-495a-af89-35115e9a1675 1175-5326 202575 Philonthus decorus (Gravenhorst) ( Figs 16–18 , 36–40 , 50–51, 57, 67) Body length: 6.60–8.00 mm (mean 7.40 mm ), female 6.60–7.98 mm (mean 7.50 mm ), male 6.90–7.40 mm (mean 7.20 mm ); body width: 3.25–3.74 mm (mean 3.4 mm ); female 3.25–3.74 mm (mean 3.4 mm ); male 3.25–3.36 mm (mean 3.31 mm ); head width: 1.77–1.95 mm (mean 1.85 mm ), female 1.80–1.95 mm (mean 1.89 mm ), female 1.77–1.89 mm (mean 1.82 mm ); pronotum width: 2.08–2.21 mm (mean 2.14 mm ), female 2.13–2.5 mm (mean 2.32 mm ), male 2.08–2.21 mm (male 2.20 mm ), about as broad as long; body stocky and strongly sclerotized. Head 1.1 x as long as wide ( Figs 16, 17 ). Antennae distinctly protruding beyond middle of elytra ( Fig. 17 ). Anterior margin of pronotum with 15–23 projections (females–9/8, 9/7, 10/9 , 12/11, 9/9, 9/10, 9/9, 10/10, 9/10, 10/10, male–8/ 8, 10/9 , 10/8 , 11/9, 8/7, 9/10 projections on sides in individuals). Wings reaching the posterior margin of 1st (morphologically 3rd) clearly visible abdominal segment ( Figs 16, 17 ). Each fore, middle, and hind tibiae with 5, 9, and 5 protuberances respectively. Tarsi of hind legs reaching to, or slightly protruding beyond, posterior margin of 4th (morphologically 6th) clearly visible, abdominal segment ( Figs 16, 17 ). Abdominal segments II and III almost equal in width, then abdomen gradually narrowed to the terminal segment of the body ( Fig. 18 ). Abdominal segments III–VIII each with a pair of setiform projections (6 pairs in total) on sides ( Figs 16–18 ). Segments III–VI with short, straight projection, segments more than 4 (4.1–4.3) x longer than projections ( Fig. 37 ); segment VII with long, slightly curved projections, segment about 1.3 x as long as projections ( Fig. 36 ); segment VIII with longest, moderately curved projections, almost as long as segment ( Fig. 39 ). All abdominal projections on sides bearing cuticular processes ( Figs 38, 40 ). Abdominal segments with microstructure as in Fig. 39 A. Female pupa with two ventral prolongations (Fig. 50); each prolongation with tiny denticle inside near apex (Fig. 50A); terminal prolongation with tiny numerous cuticular processes in apical region (Fig. 50B). The first pair of functional spiracles relatively small, similar to second, moderately elongated, without microstructure ( Figs 17, 18 , 57 ); atrophied spiracles as in Fig. 67 . FIGURES 13–18. Pupa of Philonthus debilis (13–15) and P . decorus (16–18). (13, 16) Ventral aspect, (14, 17) lateral aspect, (15, 18) dorsal aspect. FIGURES 19–40. Pupa of Gabrius osseticus (19, 20), Philonthus atratus (21–25), P . carbonarius (26–32), P . debilis (33–35) and P . decorus (36–40). (19, 21, 26, 27, 36) Lateral part of segment VII with microstructure (19A, 26A) and spine (27); (20, 22, 33, 39, 40) lateral part of segment VIII with microstructure (39A) and spine (40); (23, 24, 28, 29) lateral part of segment IV with spine (24, 29); (25, 31, 32, 34, 35, 37, 38) lateral part of segment V with microstructure (34A) and spine (32, 35, 38). FIGURES 41–51. Pupa of Bisnius fimentarius (41, 41A, B, 42), Gabrius osseticus (43, 43A, B, 44, 44A), Philonthus atratus (45, 45A, 46), P . carbonarius (47, 47A, 48, 48A), P . debilis (49, 49A, B) and P. decorus (50, 50A, B, 51). (41, 43, 44, 45, 47, 49, 50) Terminal sternite of female pupa; (42, 46, 48, 51) gonotheca or entire terminal sternite (48) of male pupa; (41A, 43A, 44A, 45A, 47A, 48A, 50A) ventral prolongations; (41B, 43B, 48A, 49B, 50B) terminal prolongation.