The sprightly little sphaerodactyl: Systematics and biogeography of the Puerto Rican dwarf geckos Sphaerodactylus (Gekkota, Sphaerodactylidae) Author Daza, Juan D. Author Pinto, Brendan J. Author Thomas, Richard Author Herrera-Martinez, Alexandra Author Scantlebury, Daniel P. Author Padilla García, Luis F. Author Balaraman, Rajesh P. Author Perry, Gad Author Gamble, Tony text Zootaxa 2019 2019-12-19 4712 2 151 201 journal article 24549 10.11646/zootaxa.4712.2.1 0c70e133-0761-4802-aa47-01182984a070 1175-5326 3586953 EBB98EFC-33EE-4831-A0EE-4C4F7032CA1A Sphaerodactylus grandisquamis guarionex comb. nov. Thomas & Schwartz, 1966 Figures 16–17 Holotype — MCZ R-81048 Type locality : “Officers’ Club Beach, Ramey Air Force Base, Puerto Rico .” [Currently called Punta Borinquen Golf and Country Club, Aguadilla , Puerto Rico .] FIGURE 16. Live specimens of Sphaerodactylus g. guarionex from Barceloneta, Puerto Rico (A, female and B, male). FIGURE17. Adult specimens of Sphaerodactylus g. guarionex from Barceloneta,Puerto Rico.A–B, Female (SHSUHerp000187), C–D male (SHSUHerp000183). Diagnosis : SVL Min/Max (12.17/ 34.2 mm ); midbody scales 36–49 (x̅ 41.3); number of dorsal body scales very reduced (around 14) compared to the other subspecies of S. grandisquamis (17–23); rounded snout scales; one internasal scale; low number of escutcheon scales (around 67, other subspecies of S. grandisquamis [73–95]); nine toe lamellae on the fourth toe; males with orange head; males throat patterned (also in S. g. mimetes and S. g. spanius ) and variable in females; the scapular patch is the largest among all members of the S. macrolepis complex, especially in females, and is usually hexagonal enclosing the ocelli. Color in life ( Fig. 16 ): Female: ground color light brown, head and tail fluctuates from orange to yellow, head pattern well defined, medial lines separated and parallel, ocelli color white. Male ground color light brown to gray, head yellow to light orange. Iris color yellow copper. Distribution : Low elevation coastal areas of northern Puerto Rico from the lowlands of El Yunque National Forest to Rincón , a transition to S. m. ateles around Mayaguez and with S. g. spanius on high elevations of the Cordillera Central.