Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini) Author Peck, Stewart B. Author Cook, Joyce text Insecta Mundi 2013 2013-06-28 2013 310 1 32 journal article 4974 10.5281/zenodo.5193714 8df27694-d6a3-43d8-8a92-a085ef706e26 1942-1354 5193714 1CC5FBEF-1373-444C-AA1C-0E80445A7B6E Cyrtusa Erichson Cyrtusa Erichson, 1842: 221 . Type species: Anisotoma subtestacea Gyllenhal, 1813: 707 ; by subsequent designation of Thomson 1859: 58. Daffner 1982: 209 ; 1983: 136 ; 1988: 292 . Caenocyrta Brown, 1937b: 172 . Type species: Amphicyllis picipennis LeConte, 1863 , by original designation (= Cyrtusa subtestacea ( Gyllenhal, 1813 )) . Synonymy by Daffner 1988: 292 . Diagnosis. Body strongly convex, oval in shape. Antenna with 10 antennomeres, with 4-antennomere club. Mandibles form semicircle when closed; right mandible with blunt tooth on anterior one-third. Ventral side of head with paired antennal grooves next to eyes. Mesosternum vertical between coxae, vertical surface not carinate. All tibiae slender; protibia with distinct tarsal groove, its outer margin strongly spinose; mesotibia slightly widened and its outer margin densely spinose; metatibia with few spines on ventral surface near outer margin. Tarsal formula 5–5– 4 in both sexes. Males distinguished by expanded pro- and mesotarsi, curved mesotibiae bearing a process on the inner apical margin, and by a toothlike expansion of the lower margin of the metafemur. Distribution. The distribution of the genus is Holarctic and it is also found in southern India ; it is questionably Afrotropical and Neotropical ( Newton 1998 ).