Systematics and distributions of the genera Cyrtusa Erichson, Ecarinosphaerula Hatch, Isoplastus Horn, Liocyrtusa Daffner, Lionothus Brown, and Zeadolopus Broun of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini) Author Peck, Stewart B. Author Cook, Joyce text Insecta Mundi 2013 2013-06-28 2013 310 1 32 journal article 4974 10.5281/zenodo.5193714 8df27694-d6a3-43d8-8a92-a085ef706e26 1942-1354 5193714 1CC5FBEF-1373-444C-AA1C-0E80445A7B6E Liocyrtusa nigriclavis ( Hlisnikovsky, 1967 ) ( Fig. 5 , 19 ) Liocyrtusa nigriclavis ( Hlisnikovsky, 1967 ) : 240; Daffner 1983: 134 ; 1988: 285 . Holotype male in HNHB (Hungarian Natural History Museum, Budapest), not seen. Type locality: Cojbalsan aimak, 820 m , 20 km SW Somon Bajanuul, Mongolia . Diagnosis. Length (pronotum + elytra) = 1.78–2.40 mm ; greatest width = 1.22–1.70 mm . Pale to dark reddish brown, mostly shiny, antennal club often darker; some individuals with microsculpture on elytra. Head moderately coarsely and densely punctate. Antennal club variable in width, more robust in larger individuals. Antennomere 8 disc-like, about half width of 9. Apical antennomere clearly narrower than 9 and 10. Sides of pronotum rounded, posterior angles obtuse. Pronotum moderately finely punctate; punctures irregularly separated, more dense laterally. Elytral strial punctures coarse, separated by less than one diameter; striae cannot be separated apically from coarse interstrial punctures. Metasternum coarsely punctate; punctation dense laterally, sparse medially. Metatibia narrow, width at apex twice width at base. Male mesotibial process short, triangular. Abdominal sternite III coarsely punctate, sternites IV–VII each with transverse row of fine punctures before apex. Median lobe of aedeagus broad, apical lobes short with rounded apices ( Fig. 5 ). Parameres ( Fig. 5 ) broad, with membranous apices extending well beyond apices of median lobe. Internal sac as in Fig. 5 . Distribution . The species is Holarctic. North American distribution ( Fig. 19 ): CANADA . ALBERTA , BRITISH COLUMBIA . MANITOBA , NUNAVUT TERRITORY , ONTARIO , QUEBEC , SASKATCHEWAN, YUKON TERRITORY . UNITED STATES . ALASKA , IDAHO , INDIANA , NORTH CAROLINA , NEVADA , UTAH , OREGON . VIRGINIA , WASHINGTON , WYOMING . Extra-limital distribution: Eastern Europe, Mongolia, Siberia. Previously recorded in North America ( Daffner 1983 , 1988 ) from: CANADA . BRITISH COLUMBIA . 25 km W Creston. 10 km W Summerland. 13 km N Nelway. ONTARIO . Chaffeys Locks. Stittsville. QUEBEC . Gatineau. UNITED STATES . VIRGINIA . Pulaski Co. , 7 mi SE Mechanicsburg. WYOMING . 34 mi E Lovell. OREGON . Clackamas Co. , Salmon River near Zigzag. WASHINGTON . Okanogan Co. , near Buzzard Lake. Seasonality. Adults have been collected in the months from June to October, with most in July to September. Bionomics. In North America, the species is mostly northern or at upper elevations in boreal forest or in tundra habitats. Adults have been collected in habitats ranging from subarctic tundra to spruce-willow and pine and pine-aspen forests to mixed hardwood forest. They have been taken mostly by flight intercept traps, but also by evening car netting, and unbaited pit-traps. New material examined. We have seen 232 specimens from the following localities. CANADA ALBERTA . Opal. MANITOBA . 100 road km SE Flin Flon. 17 km N Woodridge, Sandilands Provincial Forest. NUNAVUT TERRITORY . Kugluktuk, N67.78463 , W115.20987 . ONTARIO . 20 km SE Almonte, Middleville to White Lake. Chaffeys Locks, Queens University Biology Station. Gloucester. 15 km NW Renfrew. Hamilton. Manitoulin Island. Heckston, 20 km SE Kempville. 15 km W Ottawa, Shirleys Bay. Stittsville. L3C6 Wolford Township, 44° 52’03”N, 75°43’50” W . SASKATCHEWAN . Moose Mountain Provincial Park, Kenosee Lake. YUKON TERRITORY . Champagne, Alaska Highway km 1580. Dempster Jct., 40 km E Dawson. EMAN plot, site LMK333 Y . Haines Jct, 10 km S . Long Lake Road, site LMK25 Moose Creek, 14 km NW Stewart Crossing. Ross River. UNITED STATES . ALASKA . 11 mi S Anderson Jct, Rt. 3, mi 270, Alaska highway. Big Delta. Nenana, 13 mi NE, mile 318, Alaska Highway. Tok. IDAHO . Clark Co. , 2.8 km W rt. 115, Stoddard Campground. INDIANA . Tippecanoe Co. NORTH CAROLINA . Haywood Co. , Balsam Mt., Blue Ridge Parkway. NEVADA White Pine Co. , Snake Range, Wheeler Peak trail. UTAH . Summit Co. , Bear River Camp. WYOMING . 34 mi E Lovell, Big Horn Mountains.