Terrestrial isopods from the Oued Laou basin, north-eastern Morocco (Crustacea: Oniscidea), with descriptions of two new genera and seven new species Author Taiti, Stefano Istituto per lo Studio degli Ecosistemi, Consiglio Nazionale delle Ricerche, Florence, Italy; Author Rossano, Claudia Dipartimento di Biologia, University of Florence, Florence, Italy text Journal of Natural History 2015 2015-02-28 49 33 2067 2138 journal article 21136 10.1080/00222933.2015.1009512 c5336d61-ecae-4eb6-8592-60f43e587af8 1464-5262 3999869 DCBF3103-1463-4A32-9BC0-A4CFE8B762AE Ctenoscia minima ( Dollfus, 1892 ) ( Figures 6–8 ) Chaetophiloscia dorsalis Verhoeff, 1928 , p. 137 , Figures 34 , 37 , 38, syn. nov. Ctenoscia minima ; Vandel, 1961 , p. 256 Figure 6. Ctenoscia minima from St. 14, ♀: (A) adult specimen, dorsal view; (B) dorsal scaleseta; (C) co-ordinates of noduli laterals; (D) cephalon, dorsal view; (E) cephalon, frontal view; (F) left side of pereon showing disposition of noduli laterales; (G) pereonite 1, left side; (H) pleonites 4, 5, telson and uropods; (I) antennule. Figure 7. Ctenoscia minima from St. 14, ♀: (A) antenna; (B) left mandible; (C) right mandible; (D) maxillule; (E) maxilla; (F) maxilliped. Figure 8. Ctenoscia minima from St. 14, ♂: (A) pereopod 1; (B) pereopod 7; (C) pleopod 1 (specimen 4.5 mm long); (D) genital papilla and pleopod 1 (specimen 4.0 mm long); (E) pleopod 2; (F) pleopod 3 exopod; (G) pleopod 4 exopod; (H) pleopod 5 exopod. Material examined 2 ♂♂ , 16 ♀♀ , 1 juv. , St. 8, leg. S. Taiti and C. Rossano , 28 April 2004 ( MZUF 9464 ) ; 4 ♂♂ , 14 ♀♀ , St. 10, leg. S. Taiti and C. Rossano , 26 April 2004 ( MZUF 9465 ) ; 1 ♀ , St. 12, leg. C. Rossano , 26 April 2004 ( MZUF 9466 ) ; 12 ♂♂ , 20 ♀♀ , St. 14, Phillyrea wood, leg. S. Taiti , 27 April 2004 ( MZUF 9467 ) ; many ♂♂ and ♀♀ , St. 15, sieved ground near stream, leg. S. Taiti , 27 April 2004 ( MZUF 9468 ) ; 1 ♀ , St. 17, leg. S. Taiti , 29 April 2004 ( MZUF 9469 ) ; 1 ♀ , St. 18, cork-oak wood, leg. S. Taiti , 29 April 2004 ( MZUF 9470 ) ; 1 ♂ , 2 ♀♀ , St. 21, leg. S. Taiti and C. Rossano , 30.IX.2005 ( MZUF 9471 ) ; 2 ♀♀ , St. 24, cork-oak wood, leg. S. Taiti , 29 April 2004 ( MZUF 9472 ) . Remarks Ctenoscia minima was originally described by Dollfus (1892) from Grenada , Spain , and later recorded for several localities in Spain , including the Canary and Balearic islands , Portugal (see Schmalfuss 2003 for previous citations), and Morocco ( Vandel 1961 ). This species has been illustrated by Vandel (1946) and Rodríguez and Barrientos (1993) . The genus Ctenoscia also includes another species, C. dorsalis ( Verhoeff, 1928 ) , originally described from San Remo and Grimaldi ( Liguria , Italy ), and later recorded from several localities in mainland Spain and the Balearic Islands , Corsica , Italy including Sardinia and Sicily , Malta (see Schmalfuss 2003 for previous citations), and Tunisia ( Achouri et al. 2008b ). According to the descriptions, C. minima differs from C. dorsalis in having the distal part of the male pleopod 1 endopod bent outwards instead of straight, while all the other characters are identical in the two species. The examination of many specimens of Ctenoscia from the Oued Laou valley showed that C. dorsalis has to be considered as a junior synonym of C. minima . In fact, the distal part of the male pleopod 1 endopod is straight or bent outwards according to the size of the specimens. In the population from St. 14 (Tirinesse), 4 mm long males show the endopod bent outwards as in C. minima ( Figure 8D ), while in 4.5 mm long males this appendage is straight as in C. dorsalis ( Figure 8C ), showing that this character varies according to the age and size of the specimens. For the same reason also the distal margin of the male pleopod 1 exopod is more or less convex. Ctenoscia minima is here fully illustrated to facilitate its identification ( Figures 6–8 ). Distribution Portugal , Spain (including Canary and Balearic islands ), Corsica , Italy (including Sardinia , Sicily and surrounding islands), Malta , Morocco and Tunisia . New record for the Rif region.