Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae) Author Medrano, Miguel Author Kury, Adriano Brilhante Author Mendes, Amanda Cruz text Zoological Journal of the Linnean Society 2022 2021-08-30 195 585 672 journal article 113070 10.1093/zoolinnean/zlab043 e070d52f-d008-4f08-988a-a8b17c27418f 0024-4082 6959516 1C89556A-12CB-43B7-9B49-E02EFF1543D3 Paraprotus atroluteus Roewer, 1912: 120 . Type data: Syntypes : Ecuador . ‘ Columbien’ [Zamora- Chinchipe]: Sabanilla [–3.95843°, –79.04729°, 1800 m ], 3 ♂ 1 ♀ ( SMF RI 291 , examined by photograph) . Records: ECUADOR [ Zamora-Chinchipe ], Zamora Valley ( Roewer, 1928 ). Figure 28. Paraprotus albithorax , female holotype (SMF RII 132/2). A, dorsal view. B, lateral view. C, panoramic dorsal view. D, original label. Scale bar: 1 mm. Photos courtesy Abel Pérez. Figure 29. Paraprotus atroluteus , syntype (SMF RI 291). A, dDorsal view. B, lateral view. C, panoramic dorsal view. D, original label. Scale bars: 1 mm. Photos courtesy Abel Pérez. Diagnosis: It differs from P. albithorax by having in carapace the same colour of abdomen ( Fig. 29A ) instead of yellow spots. Abdomen with slightly yellowish coloration in lateral margins ( Fig. 29A ). Tarsal counts: 7(3), 17(3)/19(3), 9, 10. Distribution: Only known from the type locality. Eastern Cordillera real montane forest ecoregion in Zamora Chinchipe Province of Ecuador ( Fig. 23 ). Remarks: Although it was described from ‘Columbien ( Sabanilla )’ ( Fig. 28D ), the correct place is possibly ECUADOR , Zamora-Chinchipe , Sabanilla . In the description of various species, Roewer (1913 ; 1923 ) inconsistently cited Sabanilla either as being in ‘Kolumbien’ or in ‘Venezuela’. Sabanilla , in this case, probably refers to a small village called Sabanilla [–4.03333°, –79.01667°, c . 1500 m ], on the old road between Loja and Zamora ( Zamora-Chinchipe , southern Ecuador). A subsequent record ( Roewer, 1928 ) reinforces our interpretation. Although external similarity is encountered comparing the type specimens of both species of Paraprotus ( Figs 29 , 30 ), Roewer (1912: 120) made a detailed description of the coloration of P. atroluteus , remarking that dorsal scutum was mostly light rustyellow except for brown-black coloration of ocularium and ventral surface (coxae and free sternites). Then, no yellow mark is described by Roewer (1912) for P. atroluteus , which is a remarkable feature present in P. albithorax , described and illustrated by Roewer (1947: 33 , pl. 12, fig. 106) and visible in Fig. 28A.