A new integrated morpho- and molecular systematic classification of Cenozoic radiolarians (Class Polycystinea) - suprageneric taxonomy and logical nomenclatorial acts Author Suzuki, Noritoshi Author Caulet, Jean-Pierre Author Dumitrica, Paulian text Geodiversitas 2021 2021-07-08 43 15 405 573 journal article 5275 10.5252/geodiversitas2021v43a15 a8353504-9387-42cf-8d81-8ecacbe9bd90 1638-9395 5101757 urn:lsid:zoobank.org:pub:DC259A19-9B35-4B33-AD9F-44F4E1DA9983 Superfamily SPONGOPYLOIDEA Dreyer, 1889 n. stat. sensu Suzuki emend. herein Spongopylida Dreyer, 1889: 42 [as a subfamily]. DIAGNOSIS. — Test flat-disk shaped, ovoidal and spherical. Funnel-like pylome, aperture (large opening) or relevant structure is present. Very thick cortical shell develops in spherical to ovoid pattern. Flat-disc type , shows a concave central part, a peripheral inflation, and a circular depression zone between the central and peripheral zone. A margarita defined by Ogane & Suzuki (2006) is positioned in the center ( Spongopylidae ) or somewhere ( Cristallosphaeridae ). The outer around the margarita looks spongy, formed by highly dense concentric structure; however, it does not form a perfect concentrically wall (like occurs in Trematodiscidae ). Endemic form might be reported in Calcaromma due to the loss of all siliceous skeletal parts except the margarita . The presence of a thick axoflagellum might be linked to a funnel-like pylome. REMARKS This superfamily includes, Spongopylidae , Cristallosphaeridae and? Prunopylidae . This superfamily corresponds to Clade L2 of Sandin et al. (2021) based on molecular data obtained for Schizodiscus and Spongobrachiopyle (= Spongopyle in Sandin et al. , 2021 ; Spongopylidae ) and Calcaromma ( Cristallosphaeridae ). The family Prunopylidae is questionably assigned in this superfamily considering the presence of a Spongopylidae-like pylome, an aperture and a very thick outermost cortical shell, similar to that observed in Enalomelon . The diagnosis of the superfamily is based only on Spongopylidae and Cristallosphaeridae only, because little is known about the internal structure of the genera assigned in Prunopylidae .