Description Of New Species Of The Genus Pteroneta (Arachnida: Araneae: Clubionidae) From Papua New Guinea
Author
Versteirt, V.
Author
Deeleman-Reinhold, C.
Author
Baert, L.
text
Raffles Bulletin of Zoology
2008
2008-08-31
56
2
307
315
journal article
10.5281/zenodo.4508188
2345-7600
4508188
Pteroneta brevichela
,
new species
(Fig. 6)
Material examined. –
Holotype
male:
Papua New Guinea
,
Madang
[=Province],
Baiteta forest
8 Jun.1993
(
T9
, coll.
O. Missa
). Female
allotype
:
Papua New Guinea
,
Madang
[=
Province
],
Baiteta forest
8 Jul.1993
(
T9
, coll.
O. Missa
).
Paratypes
:
1 female
21 Jun.1995
(AR17),
1 female
22 Jun.1995
(AR18),
1 female
30 Jun.1995
(AR22),
1 female
30 Jun.1995
(AR25-14),
2 females
13 Jul.1995
(AR29),
2 males
17 Apr.1996
(AR43),
1 female
9 May 1996
(AR50),
1 female
7 Jun.1996
(AR56),
1 female
18 Jun.1996
(AR58),
1 female
25 Jul.1996
(AR70),
1 male
and
2 females
30 Mar.1993
(M2),
1 male
and
1 female
22 Apr.1993
(M4),
1 male
14 Jun1994
(M10),
1 female
6 Apr.1994
(
XI
).
All
from
Papua New Guinea
,
Madang
[=Province],
Baiteta forest
(
Table
1)
.
Fig. 5.
Pteroneta longichela
; A, ventral view of left male pedipalp; B, dorsolateral view of left male pedipalp, C: ventral view (epigyne). Abbrevations used: emb, embolus; spd, spermduct; ste, subtegulum; tap, tibial apohysis; ent, entrance opening; fed, fertilization duct; ind, insemination duct; lsp, lower spermathecae; usp, upper spermathecae.
Diagnosis
.
–
Males of this species are distinguished by the shorter, thick chelicerae, by the number (only one) and position of spines on the chelicerae, with bulge in mesal part of the fangs. Females of this species are characterized by the very long insemination ducts, entering caudolaterally and by the smaller upper dark spermathecae and larger more transparent lower spermathecae.
Description. –
Male: total body length: 3.0 mm; carapace length:
1.45 mm
, width:
1.1 mm
; abdomen length:
1.53 mm
and width: 1.0 mm. Eyes: PME:
0.26 mm
and PL:
0.56 mm
apart, head width:
0.74 mm
. Abdomen and legs are coloured pale yellow, carapace and chelicerae on the other hand are brownish yellow. Chelicerae with 3 promarginal teeth and 1 retrolateral tooth; only one short blunt spine on dorsal side, bunch of hairs on chelicerae mesally situated (till PMT 2). Legs: Spination: femora I-II 2*d 1p, III 2*d, IV 1d (left); tibiae I 2*v, II-III spineless, IV 1d; metatarsi I-II spineless, III 102d 2*v + circle of 11 (barbed) spines, IV 202d 1p 102v + retrolateral bunch of 11 small spines + 1 ventrally situated. Measurements: Fe I:
0.86 mm
, Pa+Ti I:
1.2 mm
, Mt I:
0.53 mm
, Ta I:
0.36 mm
; Fe II:
1.08 mm
, Pa+Ti II:
1.54 mm
, Mt II:
0.56 mm
, Ta II:
0.55 mm
; Fe III:
0.59 mm
, Pa+Ti III:
0.81 mm
, Mt III:
0.58 mm
, Ta III:
0.26 mm
; Fe IV:
0.95 mm
, Pa+Ti IV:
1.18 mm
, Mt IV:
0.94 mm
, Ta IV:
0.3 mm
.
Male pedipalp (see Fig. 6 A–B): Tibial apophysis similar to
P. longichela
with exception of the tip, which is much sharper; tibia without any spines. Sperm duct originating from larger structure, going distally, making a curve of 180° and ending in a sharp beak like embolus. Tegulum longer than wide with additional small, sharp ridged appendage (retrolateral flange), distally, near embolus basis. Cymbium with 2 apical spines on ventral side.
Female: total body length:
3.5 mm
; carapace length:
1.4 mm
, width:
1.03 mm
; abdomen length:
1.98 mm
and width:
1.15 mm
. Eyes: PME:
0.2 mm
apart, PL:
0.44 mm
and head width:
0.6 mm
. Sometimes very pale yellow spiders (abdomen and carapace), chelicerae and fangs are slightly darker. Chelicerae with 6 very small promarginal teeth and 6 dot like retromarginal teeth. Legs: Spination: femora I-IV 2*d; tibiae I-III spineless, IV 1p 1r, metatarsi I 200v, II (10- 01-0)v, III 102d 1v + circle of 10 (barbed) spines, IV 202d 102v + retrolateral bunch of 9 small spines. Measurements: Fe I:
0.64 mm
, Pa+Ti I:
0.98 mm
, Mt I:
0.39 mm
, Ta I:
0.3 mm
; Fe II:
0.94 mm
, Pa+Ti II:
1.23 mm
, Mt II:
0.36 mm
, Ta II:
0.55 mm
; Fe III:
0.56 mm
, Pa+Ti III:
0.61 mm
, Mt III:
0.46 mm
, Ta III:
0.23 mm
; Fe IV:
0.7 mm
, Pa+Ti IV:
1.15 mm
, Mt IV:
0.76 mm
, Ta IV:
0.33 mm
.
Epigyne (see Fig. 6 C): circular and smaller upper spermathecae and two larger and circular lower spermathecae. Long insemination ducts originating from caudo-laterally situated entrance openings; making a loop of 180° ending in the proximal part of the lower spermathecae. Short and wide fertilization duct originating from upper spermathecae.
Etymology. –
After the shorter male chelicerae, by which it differs from the previous species.
Remarks. –
Pteroneta longichela
and
P. brevichela
are very alike in habitus (as do most
Pteroneta
species) but differs in a number of somatic characteristics. The most important ones in male individuals are the length of the chelicerae (and number of spines) and the number of promarginal retromarginal teeth. The pedipalps of both species are slightly different, whilst the epigynes can easily be distinguished (see Figs. 5–6). Preliminary cladistic analyses show that they are closely related but differ enough to be placed in different branches.