Taxonomy and anatomy of Amphiboloidea (Gastropoda: Heterobranchia: Archaeopulmonata)
Author
Golding, Rosemary E.
Author
Ponder, Winston F.
Author
Byrne, Maria
text
Zootaxa
2007
1476
1
50
journal article
10.5281/zenodo.176773
1a670eb6-5f50-42d6-ace0-3e52fd4f4c17
1175-5326
176773
Genus
Naranjia
gen. nov.
Type
species
Salinator
cf.
swatowensis
Yen, 1939
(
Hong Kong
form).
Etymology
: Derived from
narangi
(Hindi—orange), referring to the brownish-orange colour of the shell.
Diagnosis:
Shell diameter to
7 mm
, fragile, brownish-orange, smooth; last whorl inflated, sometimes with spiral ridge on rounded shoulder, spire short; umbilicus intermediate width, lacking umbilical fold; aperture with flattened, indented sinus. Operculum thin, corneous, pale yellow with eccentric, paucispiral nucleus, inner surface lacking elaboration around nucleus. Head unpigmented in formalin-preserved specimens. Mantle cavity with opposed ciliary tracts restricted to exhalant canal; hypobranchial gland containing large, mucus-filled, chambers. Radula with central tooth, single lateral tooth, approximately 25 marginal teeth on each side of radula; central tooth with 13–17 cusps, mesocone enlarged, adjacent cusps bearing numerous smaller cusps, other cusps decreasing in length towards outer edge of tooth; lateral teeth with u-shaped gap between innermost and adjacent cusps; marginal teeth elongate, narrow. Central nervous system euthyneurous, cerebral and pleural ganglia fused; presence of subcerebral and parapedal commissures not confirmed; visceral loop concentrated, parietal ganglia positioned asymmetrically, accessory ganglion lacking. Seminal vesicle diverticulum from upper hermaphrodite duct, seminal receptacle black pigmented. Spermovipositor folded across aperture of spermoviduct, with half-spiral plate fused to distal spermovipositor sheath; numerous long, finger-shaped papillae covering distal spermovipositor. Spermovipositor sheath with enlarged pocket on left of spermovipositor, forming ‘accessory bursa’ with narrow aperture. Prostatic gland blind-ending, tubular, uniform for entire length. Egg mass unknown.
Remarks:
The
type
species of this genus is intended to be the
Hong Kong
taxon described below, not
Salinator swatowensis
Yen 1939
, if it proves to be distinct.
Salinator swatowensis
was originally included in
Salinator
based on shell characters (
Yen 1939
). However, anatomically, the
Hong Kong
material attributed to
N.
cf.
swatowensis
is clearly separable from species of
Salinator
. The central and lateral radular teeth are distinctive and the spermovipositor is folded and bears long, unchitinised papillae. The spermovipositor sheath has a unique enlarged pocket and the hypobranchial gland has large mucus chambers which are not present in
Salinator
.