Pelagic tunicates (Appendicularia and Thaliacea) of Sri Lanka: two first records with an annotated checklist
Author
Karunarathne, Krishan D.
0000-0003-1780-9749
Department of Aquaculture and Fisheries, Faculty of Livestock, Fisheries and Nutrition, Wayamba University of Sri Lanka, Makandura, Gonawila, 60170, Sri Lanka. & krishankarunarathne @ gmail. com; krishan. dk @ wyb. ac. lk; https: // orcid. org / 0000 - 0003 - 1780 - 9749
krishankarunarathne@gmail.com
Author
Croos, M. D. S. T. De
0000-0003-4449-6573
Department of Aquaculture and Fisheries, Faculty of Livestock, Fisheries and Nutrition, Wayamba University of Sri Lanka, Makandura, Gonawila, 60170, Sri Lanka. & dileepa _ dc @ yahoo. com; dileepad @ wyb. ac. lk; https: // orcid. org / 0000 - 0003 - 4449 - 6573
dileepa_dc@yahoo.com
text
Zootaxa
2021
2021-11-11
5067
3
352
376
journal article
3575
10.11646/zootaxa.5067.3.2
9b8d33b9-c9df-4f9d-abeb-de1c7b84f3e0
1175-5326
5681899
AF3DF45F-B248-4BDD-904F-F55D735DC867
Thalia sibogae
Van Soest, 1973
Thalia sibogae
Van Soest 1973: 204
, 205, Fig. 8; 1998: 235, Table 14.1.
Kim
et al
. 2011: 147
, Fig. 5.
Purushothaman
et al
. 2017
.
Ishak
et al
. 2018: 454
, 459–465,
Figs. 4a–e
, 5.
Material examined.
One solitary zooid (
MDAFWU 2018
/85),
St
.133,
January 2018
(
Fig. 2D–F
)
.
Description.
Solitary zooid: Transparent body bulky rounded with a very thick, smooth test, with two long (about one third of body length), tapering processes on the posterior end (
Fig. 2D–F
); length without processes
7 mm
. Atrial palps fairly small and bifurcate (
Fig. 2F
); lateral projections very small, simple and not bifurcated (
Fig. 2D, F
); medioventral projections are small and of unequal length, the anterior ones are the smallest (
Fig. 2E
). Six body muscles broad; M I narrowly interrupted dorsally, M II–IV converged mid-dorsally, M V and M VI near but do not touch dorsally; M II–V in complete rings, M VI interrupted ventrally (
Fig. 2D
). Totally (M I–VI) 60 muscle fibers. The nucleus (stomach + intestine) is close to the middle posterior projection (
Fig. 2D–F
).
FIGURE 2.
Firstly reported pelagic tunicates to Sri Lanka in “
Waya-jel
-Survey” (
A–F
): a live colony of
Pyrostremma spinosum
near to the sea surface (
A
&
B
), and an illustration of its blastozooid (
C
);
Thalia sibogae
solitary zooid in preserved form (
D
), ventral view of medioventral projections (
E
), and an illustration of its posterior part (
F
).
a
—anus;
ap
—atrial palp;
dmp
— dorsal medioventral projection;
e
—endostyle;
es
—excurrent/buccal siphon;
g
—ganglion;
is
—incurrent/atrial siphon;
lp
—lateral projection;
n
—nucleus;
o
—ovary;
pb
—pharyngeal basket;
pp
—posterior projection;
sl
—stolon;
sm
—stomach;
t
—testis;
vmp
—ventral medioventral projection. Scales: E = 0.5 mm; C, D, F = 1 mm; A & B = 0.4 m. Images A & B by Franco Banfi from off Mirissa, Sri Lanka
Aggregate zooid: Not collected.
Global distribution.
The Central Indo-Pacific (
Van Soest 1973
,
1998
;
van der Land 2008
); Korean waters (
Kim
et al
. 2011
); the east coast of Peninsular
Malaysia
(
Ishak
et al
. 2018
).
Type
locality: Pulu Sanguisapio, Tawi-Tawi Island, Sulu archipelago (
Siboga
-Expedition
St
.
93, 05°N
,
120°E
).
Remarks.
Although
Van Soest (1973)
did not observe lateral projections on the
type
material of
T. sibogae
, tiny lateral projections may exist with solitary zooids of
T. sibogae
according to
Kim
et al
. (2011)
and
Ishak
et al
. (2018)
. Therefore, the
Sri
Lankan specimen resembles the specimens of
Kim
et al
. (2011)
and
Ishak
et al
. (2018)
. The total number of muscle fibres (M I–VI) in solitary zooids of
Thalia democratica
(
Forskål, 1775
)
shows a clinal variation, decreasing from higher to lower latitudes (
Van Soest 1975b
); and there is a possibility to be such a variation in
T. sibogae
, but data is not enough in the literature for comparison.
Local name:
Indi Gaduwa
(in Sinhalese) [note: usually fishers use this term for tiny salps, while the
Pegea
-like larger salps are called ‘
Rata-indi Gadu
’ in Sinhalese and ‘
Pereechcham Katti
’ in Tamil].
Local importance: None [but,
Pegea
-like larger salps regularly clog into fishing nets (authors’ observations)].