A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae) Author Díaz, Florentina Author Spinelli, Gustavo R. Author Ronderos, María M. text Journal of Natural History 2010 2010-11-05 44 45 - 46 2825 2849 http://dx.doi.org/10.1080/00222933.2010.512424 journal article 10.1080/00222933.2010.512424 1464-5262 5212142 Dasyhelea gargola Díaz and Spinelli sp. nov. ( Figures 2 , 4 ) Diagnosis The only species of the patagonica group with the posteromedian margin of the male sternite 9 with a deep rounded excavation, and the posterolateral margins of the excavation broad and truncate. The female subgenital plate is rectangular, with irregular anterior margin. Male. Similar to female with usual sexual differences. Flagellum as in Figure 4A . Palpus ( Figure 4B ) with third segment bearing scattered sensilla; PR 4.23, fifth segment missing. Scutellum with 18 long setae and 18 shorter ones. Wing ( Figure 4C ) length 1.74 mm , width 0.50 mm , CR 0.51. Genitalia ( Figure 4D ): tergite 9 tapering distally, extending to level of apex of gonocoxite, apicolateral process slender, with apical seta; cercus with three or four setae; sternite 9 ( Figure 4E ) 0.5 times longer than greatest width, posteromedian margin with deep, rounded anteromesal excavation, posterolateral margins of excavation broad, truncate. Gonocoxite stout, 1.9 times longer than greatest width, with short, blunt anteromedian process; gonostylus slender, as long as gonocoxite, narrow base, nearly straight, tip pointed. Parameres ( Figure 4F ) with basal arms stout, with conspicuous laterodistal, pointed processes; anteromesad projection short, blunt, fused to posteromedian projection, the latter slender, slightly produced beyond distal portion of aedeagus body, with lightly recurved tip. Aedeagus well sclerotized, 0.75 times longer than greatest width, basal arch extending to 0.25 of total length; basal arms stout, short, recurved; body stout with triangular pointed, divergent posterolateral projections; posterolateral arms convergent, tips recurved mesad. Figure 4. Dasyhelea gargola Díaz and Spinelli sp. nov. : (A–F) male, (G–L) female. (A) Flagellum. (B) Right palpus. (C) Wing. (D) Genitalia (ventral view). (E) Sternite 9. (F) Parameres. (G) Flagellum. (H) Clypeus. (I) Right palpus. (J) Wing. (K) Subgenital plate. (L) Spermathecae. Scale bars 0.05 mm. Female. Head dark brown. Eyes contiguous by width of two to three ommatidia. Antenna with flagellum ( Figure 4G ) dark brown; AR 1.13 (1.09–1.17, n = 2). Clypeus ( Figure 4H ) with 14–17 setae. Palpus ( Figure 4I ) dark brown; third segment with four sub-basal capitate sensillae; PR 3.82 (3.71–3.93, n = 2). Thorax. Scutum dark brown, scutellum with 19–20 long setae and 15–16 shorter ones. Legs dark brown, including tarsi; hind tibial comb with five spines; prothoracic TR 1.88, mesothoracic TR 1.90, metathoracic TR 1.87 (1.84–1.90, n = 2). Wing ( Figure 4J ), length 1.46 mm ( 1.44–1.48 mm , n = 2), width 0.58 mm ( 0.56– 0.60 mm , n = 2), CR 0.52 ( n = 2); membrane slightly infuscated, densely covered with macrotrichiae; second radial cell open; cubital fork at same level of anterior portion of second radial cell. Halter dark brown. Abdomen. Dark brown. Subgenital plate ( Figure 4K ) rectangular, anterior margin irregular; posterolateral arms stout, posteromedian projection stout, inner margin straight distally. Two ovoid spermathecae ( Figure 4L ), subequal, with straight necks, measuring 0.048 by 0.042 mm and 0.044 by 0.038 mm , necks 0.004 mm . Distribution Argentina , only known from its type locality . Type material Holotype male, Argentina , Rio Negro , Parque Nacional Nahuel Huapi , cerro Catedral , laguna Toncek , 41 ◦ 11 ′ 52.2 ′′ S , 71 ◦ 29 ′ 31.2 ′′ W , 1770 m , 20 February 2007 , C. Cazorla – M. Donato , red . Paratypes , 2 females , same data ( MLP ) . Etymology The specimens herein described as a new species were collected in the laguna Toncek located in the cerro Catedral. The specific epithet refers to the gargoyles that are common sculptures of Cathedrals. Discussion The male genitalia of this species are similar to those of D . ona , especially by the aedeagus stout body and the triangular pointed, divergent posterolateral projections. However, it can be distinguished from D . ona by several other characteristics that may be found in the discussion under the description of the later species.