The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution Author Rosa, Cecilia Lozano De La 36CFEA67-71C5-4F3C-97F8-B57E43C811CC Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela. lozanoceci@gmail.com Author Cupello, Mario BDB03C18-7095-4EAA-8BDD-03CB4F79676D Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A & M University, College Station, TX, 77843, USA. mcupello@hotmail.com Author Vaz-De-Mello, Fernando Z. 2BC093C2-3E2B-466B-B31C-9F3D9FA844BC Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil. vazdemello@gmail.com text European Journal of Taxonomy 2024 2024-10-11 959 1 1 272 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401 journal article 10.5852/ejt.2024.959.2677 2118-9773 13945464 30872C13-516F-42FA-AFA7-30ADC6BF1BAF Dichotomius satanas (Harold, 1867) Fig. 43A Pinotus satanas Harold, 1867d: 98 (original description). Type locality: Colombia . Name-bearing type: syntypes (MNHN), examined by FZVM. Pinotus satanas Harold 1880: 24 (distribution for Colombia ). — Blackwelder 1944: 208 (list, distribution). Gillet 1911b: 62 (catalogue). Dichotomius satanas Halffter & Matthews 1966: 83 (comments). — Howden & Young 1981: 126 (redescription, distribution). — Deloya 1992: 2 (list). — Halffter et al. 1992: 149 (ecology). — Kohlmann & Solís 1997: 345 , 367 (key, redescription, cited for Panama ). — Maes 1998: 655 (list for Nicaragua ). — Escobar 2000: 208 (checklist for Colombia ). — Medina et al. 2001: 138 (list, cited for Colombia ); 2002: 182–185 (ecology). — Estrada & Coates-Estrada 2002: 1911 (ecology). — Ramírez & Locarno 2004: 61–63 (list). — Kohlmann et al. 2007: 29 (checklist). — Pulido-Herrera et al. 2007: 307 (cited for Andean region of Colombia ). — Giraldo et al. 2011: 118 (comments). — Cultid-Medina et al. 2012: 48 (guide). — Delgado et al. 2012: 327 (list). — Noriega et al. 2012: 47 (list for Antioquia , Colombia ). — Solís & Kohlmann 2012: 6 (checklist for Costa Rica ). — Noriega & Navarrete-Heredia 2013: 191 (comments). — Pardo-Locarno & Camero 2014: 210‒212 , 216 (list for Chocó, Colombia ). — Torres et al. 2017: 48–49 , 67 (cited for Boyaca , Colombia ). — Nunes & Vaz-de-Mello 2019: 2239 (comments). — Pardo-Díaz et al. 2019: 2 , 5‒12, 14 (comments). Material examined VENEZUELA Táchira 1 ♂ ; Vía Chorro del Indio ; Oct. 1984 ; Joffre B. leg.; CEMT 1♀ ; same data as for preceding except for the date; Oct. 1984 ; CEMT . Distribution Mexico , Belize , Guatemala , Nicaragua , Costa Rica , Panama , Colombia , Venezuela ( new country record ), Ecuador , and Peru . Subregion of Venezuela Andes mountains. Literature record Not reported for Venezuela .