Fossil butterflies, calibration points and the molecular clock (Lepidoptera: Papilionoidea)
Author
Jong, Rienk De
text
Zootaxa
2017
4270
1
1
63
journal article
32975
10.5281/zenodo.583183
6c479acc-8b18-4f0b-a6e5-85bcd6d7b6b7
1175-5326
583183
2D00AFF5-4FE2-4EC1-A328-C8670CFB8D6D
candiope
.
Charaxes candiope
(Godart, 1824)
Nymphalidae
:
Charaxinae
.
Pleistocene copal from
East Africa
(
Zanzibar
), but see the comments below.
Copal is half-fossilized resin, an intermediate stage of polymerization and hardening.
Depository: unknown.
Published figures: A picture of the fossil has not been published. Good pictures of the extant species can be found in many books on African butterflies.
This specimen was listed by
Skalski (1976)
without further information. It is a widespread extant species, occurring over most of sub-Saharan Africa. It is highly remarkable that dripping resin could trap a member of this genus of strong and fast-flying species, unless it was already dead or crippled.
The fossil is listed by
Sohn
et al
. (2012)
as originating from "Tanzania: Zanzibar Island".
However, as explained by
Evers (1907)
, the so-called Zanzibar-Copal does not originate from the island, but from the costal area of
Tanzania
between Bagamoyo to
Lindi
. The name "Zanzibar-Copal" may refer to the time when there was a sultanate of
Zanzibar
that covered the island as well as a large part of present
Tanzania
.
According to
Evers (1907)
, the Zanzibar-Copal is only two- to three thousand years old. It is too young to be of interest for calibration purposes, and since there is no way to check Skalski's record, it seems better to dismiss its relevance.