AlveOderes gen. nov., the earliest fossil of Bothrideridae from mid-Cretaceous Burmese amber (Coleoptera: Coccinelloidea) Author Li, Yan-Da State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences. Nanjing 210008, China. & Bristol Palaeobiology Group, School of Earth Sciences, University of Bristol. Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, United Kingdom. cycai@nigpas.ac Author Cai, Chen-Yang State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences. Nanjing 210008, China. text Zoologia 2024 e 23084 2024-06-07 41 1 10 http://dx.doi.org/10.1590/s1984-4689.v41.e23084 journal article 10.1590/S1984-4689.v41.e23084 1984-4689 13176683 Alveoderes yamamotoi Li & Cai sp. nov. Figs 1–13 https://zoobank.org/ 76A6CB73-4A51-4939-AE5C-A97BF5AE411B Material. Holotype , NIGP203385 . Locality and horizon. Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District, Kachin State , Myanmar ; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian ( Shi et al. 2012 , Mao et al. 2018 ). Diagnosis. As for the genus. Description. Body elongate, about 1.5 mm long, 0.4 mm wide; surface with scattered setae. Head declined at base. Compound eyes with short interfacetal setae. Antennae likely 11-segmented; antennomere 1 robust; antennomere 2 wider than 3; antennomeres 3–8 submoniliform to subfiliform; antennomeres 9–11 forming loose club; antennomere 11 conical. Pronotal disc about 1.1 times as long as wide, widest anteriorly; surface with median longitudinal groove, about 0.8 times as long as pronotal length, anteriorly acute, gradually widened backwards, posteriorly open, posterolaterally bordered by distinct posteriorly directed projections; anteri- or pronotal angles each with a distinct oval cavity (possibly a glandular opening). Procoxae subcontiguous (status of coxal cavities unknown). Figures 3–4. General habitus of Alveoderes yamamotoi Li & Cai gen. et sp. nov. , holotype, NIGP203385, under confocal microscopy: (3) dorsal view; (4) ventral view. Scale bars: 500 Μm. Scutellar shield elongated oval, anteriorly emarginate. Elytra elongate; surface with indistinct rows of fine punctures; one distinct longitudinal ridge present at base of elytron, anteriorly opposite to respective projection on pronotum, shorter than 1/4 elytral length, gradually weakened posteriorly; elytral humeri distinctly ridged; epipleura narrow. Mesoventrite with median carina and paired parallel carinae continuous with edges of mesoventral process. Mesocoxal cavities narrowly separated. Metaventrite anteromedially slightly tumid, without clear median discrimen; katepisternal suture absent. Metacoxae suboval, moderately separated, laterally separated from elytra by metanepisternum. Legs well-developed, slender. Trochanters relatively reduced and somewhat concealed. Tibiae with two or three weak spines along outer margin; protibiae with strongly unequal spurs; one being enlarged and strongly curved; meso- and metatibiae with paired equal spurs. Tarsi all 4-segmented; tarsomeres simple, unlobed. Pretarsal claws simple. Abdomen with five ventrites. Ratio of ventrite lengths along middle: 5.2:2.0:1.0:1.1:2.0. Ventrite 1 with intercoxal process apically broadly angulate; postcoxal lines absent. Etymology. The species is named after the coleopterist Dr. Shûhei Yamamoto. Remarks. The antennomere 11 of the left antenna is not preserved in the specimen. The prosternal process cannot be observed, and the state of procoxal cavities cannot be confidently determined.