Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae) Author Kataev, Boris M. Author Liang, Hongbin text Zootaxa 2015 3920 1 1 39 journal article 10.11646/zootaxa.3920.1.1 7289c86d-0ed1-4e93-b7b3-bc596042d854 1175-5326 287886 025F9918-F9F6-4049-B7CD-5AC632094F5E The calceatus species group Platus Motschulsky, 1844 : 197 . Type species: Harpalus calcitrapus Motschulsky, 1844 (= H. calceatus Duftschmid, 1812 ), designated by Noonan (1976: 33) . Pardileus des Gozis, 1882 : 289. Type species: Carabus calceatus Duftschmid, 1812 , by monotypy. Neopardileus Habu, 1954 : 283 (as a subgenus of Ophonus ). Type species: Ophonus itoshimanus Habu, 1954 (= H. calceatus Duftschmid, 1812 ), by original designation. Diagnosis . Pronotal basal margin bordered (sometimes border interrupted or indistinct medially or laterally). Elytra without row of discal pores on intervals 1, 3, 5 and 7. Protibia with three (rarely two) ventroapical spines arranged in a transverse row. Tarsomere 5 with spines ventrally (in addition to thin setae and hairs) (Fig. 10). Composition . This species group includes only the Palaearctic H. calceatus .