A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215 Author Van Bakel, Barry W. M. Author Guinot, Danièle Author Artal, Pedro Author Fraaije, René H. B. Author Jagt, John W. M. text Zootaxa 2012 2012-02-29 3215 1 1 216 https://biotaxa.org/Zootaxa/article/view/zootaxa.3215.1.1 journal article 10.11646/zootaxa.3215.1.1 1175­5334 5248640 B20CD4A6-D150-4CCF-931F-ED6D7EA54E8C Subfamily Cyrtorhininae Guinot, 1993 Cyrtorhininae Guinot, 1993b: 1330 . Type genus. Cyrtorhina Monod, 1956 . Genera included. Antonioranina n. gen. ; Cyrtorhina Monod, 1956 . Diagnosis. Carapace ovate, variously wide, convex in both directions. Cervical groove obsolete, branchiocardiac groove indistinct. Posterolateral margins straight, anterolateral margins rounded; margins rimmed or lined by granules. Anterolateral margin with 2, 3 teeth, subhepatic spine may be present. Cuticle microstructure diverse: upright nodes, fungiform nodes, inclined nodes, pits. Front short, widely triangular, weakly to strongly sulcate, with sharp, anteriorly directed tooth on each side. Supraorbital margin with 2 fissures, inner fissure proportionally short, outer fissure longer, separating extraorbital teeth; outer extraorbital tooth robust, falciform. Orbits small; eyestalk short, of 3 elements: proximal article reduced, only ventrally visible; second article triangular, elongated; third dorsally exposed, calcified, strongly arched, with fungiform granules; cornea small. Antennary fossae situated posterior to antennulary fossae. Antennule deeply inserted, completely hidden by antenna, basal article notched distally, following articles much smaller; flagellum short. Proepistome distinct. Antennae massive, meeting medially, with external surface granular; articles 1, 2 fused, urinal opening at inner angle; article 3 expanded, with spatulate dorsal lobe, pointed ventral lobe; article 4 with salient anterodorsal angle; article 5 short; flagellum developed. Epistome triangular, medially raised, keeled. Subantennary lobe of pterygostome not delimited. Mxp3 operculiform; endopod long, blunt anteriorly, merus shorter than ischium; palp short, concealed. Exopod short, wider than endopod, anteriorly pointed. Pterygostome granular, ridged, with rows of dense setae, completely setiferous posteriorly. Exposed pleurites 5–7 forming large, unexcavated plate; branchiostegite developed, high, overhanging exposed pleurites, limited by P5 apposed along branchiostegal margins. Sternum/pterygostome junction very narrow, recessed. Sternum/exposed peurite connections large between P1, P2, narrower between P2, P3. Thoracic sternum narrow all along, thus P1‒P5 coxae approximated. A strong deflection at level of sternite 8. Sternite 1, 2 in a lower level, concealed; sternite 3 exposed, as small, V-shaped, horizontally extended strip; sternite 4 narrow, not laterally expanded except the slender, recessed anterolateral extensions forming sternum/pterygostome junction; sternites 4, 5 expanded laterally between P1, P2; sternites 6, 7 conspicuously narrow, linear; sternite 8 perpendicular to sternite 7. Medial line along sternites 5–8. Spermathecal apertures recessed, small, contiguous. Chelipeds homochelous, homodontous. Basis-ischium short, immoveably fused with massive merus. Propodus longer than wide, narrow, upper, lower margins smooth, except for acute superodistal spine; fingers long, acicular, crossing at acute tips; dactylus deflected, proximally with long acute spine, prehensile border unarmed; fixed finger only slightly deflected, prehensile border with 3 acute spines proximally. P2–P4 moderately stout; carpus short, modified. P2–P4 dactylus spatulate on P2, sickle shaped on P3, P4. P5 dorsal, moderately reduced, granular, lying along posterolateral carapace margin, adapted to branchiostegal edge; dactyl narrow, elongated. Abdomen proportionally small but rather wide in male, freely articulated, 6 articles plus telson, incompletely folded, first 4 articles dorsal, first somite as wide as posterior carapace margin. Female abdomen only slightly enlarged, thus sexual dimorphism indistinct. Uropods or sockets absent. Large posterior branchial orifices present.