Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China Author Cai, Yan-Peng Author Zhou, Hong-Zhang text Zootaxa 2015 3990 2 151 196 journal article 10.11646/zootaxa.3990.2.1 2dd793f4-d2f7-46c2-abbf-9952384cb55b 1175-5326 245442 C3F0F9F4-2879-402C-8504-D12810018E81 Quedius ( Raphirus ) muscicola Cameron , 1932 ( Figs. 15, 15-1 ) Cameron , 1932 : 295 ( Quedius ; subgenus Raphirus ; Type locality: India , Ghum district); Scheerpeltz, 1933 : 1450 (catalog); Scheerpeltz, 1965 : 112 ( Myanmar ); Smetana, 1988 : 253 ( lectotype designation; redescription; India , Myanmar , Nepal ); Smetana, 1992 : 5 ( Nepal ); Smetana, 1997 : 131 ( China (Yunnan)); Herman, 2001 : 3214 (world catalog); Smetana, 2004 : 667 (catalog for Palaearctic region); Smetana, 2012a : 44 (redescription; China (Guizhou, Hubei, Shaanxi, Sichuan, Yunnan)). Quedius ( Raphirus ) dohertyi Cameron , 1932 : 297 ( Quedius ; subgenus Raphirus ; Type locality: Myanmar , Ruby Mines); Scheerpeltz, 1933 : 1438 (catalog); Smetana, 1988 : 253 (synonym of muscicola ); Herman, 2001 : 3214 (world catalog); Smetana, 2004 : 667 (catalog for Palaearctic region). Quedius ( Raphirus ) heterogaster Cameron , 1944 : 14 ( Quedius ; subgenus Raphirus ; Type locality: India , Chakrata district, Kanasar); Smetana, 1988 : 253 ( lectotype designation; synonym of muscicola ); Herman, 2001 : 3215 (world catalog); Smetana, 2004 : 667 (catalog for Palaearctic region). Material examined. CHINA : Sichuan Province: 1 ♂ , Wan County, Wangerbao Nature Reserve, 28. V. 1994 , coll. Xingke Yang; 1 ♂ , Dujiangyan City, Qingcheng Mountain, 4. VI. 1997 , coll. Haisheng Zhou; 4 ♂♂, 9 ♀, Baoxing County, Fengtongzhai Nature Reserve, 6–7. VI. 1997 , coll. Haisheng Zhou; 1 ♂ , Baoxing County, Yongfu, Pujigou, 2300 m , 11–14. VIII. 2003 , coll. Xiaodong Yu; 1 ♂ , Wolong National Nature Reserve, Wulidun, 2460 m , 5–8. VIII. 2004 , coll. Xiaodong Yu; 1 ♀, Wolong National Nature Reserve, Wulidun, 2270 m , 22–25. IX. 2004 , coll. Xiaodong Yu; 1 ♂ , 1 ♀, Dujiangyan City, Qingcheng Mountain, 21. VIII. 2005 , coll. Yejun Zhang; 1 ♀, Dujiangyan City, Qingcheng Mountain, 23. VIII. 2005 , coll. Xiaoyan Li; 1 ♂ , Dujiangyan City, Longchi Mountain, 26. VIII. 2005 , coll. Caiyun Zhao; 1 ♂ , 3 ♀♀, Dujiangyan City, Longchi Mountain, 1800 m , 26. VIII. 2005 , coll. Xiaoyan Li; 1 ♂ , Emeishan City, Emei Mountain, 28. VIII. 2005 , coll. Caiyun Zhao; 2 ♀♀, Emeishan City, Emei Mountain, 28. VIII. 2005 , coll. Yejun Zhang; 2 ♀♀, Emeishan City, Emei Mountain, 1200 m , 29. VIII. 2005 , coll. Xiaoyan Li; 1 ♀, Baoxing County, Fengtongzhai Nature Reserve, Dashuigou, 1500 m , 2. IX. 2005 , coll. Hongzhang Zhou and Caiyun Zhao; 1 ♂ , 2 ♀♀, Baoxing County, Fengtongzhai Nature Reserve, Dashuigou, 1500 m , 3. IX. 2005 , coll. Caiyun Zhao; 4 ♂♂, 2 ♀♀, Baoxing County, Fengtongzhai Nature Reserve, Dashuigou, 1500 m , 2–3. IX. 2005 , coll. Xiaoyan Li; 1 ♂ , 1 ♀, Lushan County, Longmen Village, 15. X. 2005 , coll. Haifeng Chen. Measurements. BL = 6.9 mm , BW = 1.4 mm , HL/PL/EL = 1.00: 1.10: 1.30, HW/PW/EW/AW = 1.00: 1.11: 1.42: 1.27, HW/HL = 1.02, HEL/HTL = 8.96, PW/PL = 1.03, EW/EL = 1.11, ESL/EL = 0.55. Distribution. Quedius muscicola Cameron is known from Myanmar , and from the mountain areas from the Himalayan range ( India , Nepal ) to southwestern China (Gansu, Guizhou, Hubei, Shaanxi, Sichuan and Yunnan Province), at altitudes from 650 m to 2800 m . Many of the examined specimens were collected in bushes, stacks of withered grass, leaf litter, or at the edges of creeks.