Pentamera fonsecae n. sp. a new species of sea cucumber from the Pacific coast of Costa Rica (Holothuroidea: Dendrochirotida: Thyonidae)
Author
Solis-Marin, Francisco Alonso
Colección Nacional de Equinodermos “ Dra. Ma. Elena Caso Muñoz ”, Laboratorio de Sistemática y Ecología de Equinodermos, Instituto de Ciencias del Mar y Limnología (ICML), Universidad Nacional Autónoma de México (UNAM), Ciudad de México, C. P. 04510, México. fasolis @ cmarl. unam. mx; http: // orcid. org / 0000 - 0001 - 5729 - 3316
fasolis@cmarl.unam.mx
Author
Alvarado, Juan Jose
Centro de Investigación en Ciencias del Mar y Limnología (CIMAR), Ciudad de la Investigación, Universidad de Costa Rica, San Pedro, 11501 - 2060 San José, Costa Rica. juan. alvarado @ ucr. ac. cr http: // orcid. org / 0000 - 0002 - 2620 - 9115
Author
Conejeros-Vargas, Carlos Andres
Posgrado en Ciencias del Mar y Limnología, UNAM, Av. Ciudad Universitaria 3000, C. P. 04510, Coyoacán, Ciudad de México, México. conejeros @ ciencias. unam. mx; http: // orcid. org / 0000 - 0003 - 0148 - 0475
conejeros@ciencias.unam.mx
Author
Caballero-Ochoa, Andrea Alejandra
Facultad de Ciencias, UNAM, Circuito exterior s / n, Ciudad de México, C. P. 04510, México. & Posgrado en Ciencias Biológicas, Universidad Nacional Autónoma de México, Ciudad de México, México.
text
Zootaxa
2020
2020-11-16
4878
3
581
588
journal article
8047
10.11646/zootaxa.4878.3.9
0df1cd40-d273-4ddb-9fd7-dca7fc392a8b
1175-5326
4425512
F91B145A-D220-4634-B13B-1B66514333BF
Key to the
Pentamera
species from the Eastern Pacific (modified from
Lambert 1998
;
Prata & Christoffersen 2016
).
1 Tube feet with supporting tables......................................................................... 2
- Tube feet lacking supporting tables.........................................
Pentamera chierchiae
(
Ludwig, 1887
)
2 Cylindrical form. Oval to elongated tables with four central holes and a short two-pillared spire table. Tube feet supporting tables with a tall spire.................................................................................. 3
- U-shaped to curved body form, podia in five bands. Circular to triangular tables, more than four central holes and a short to tall two-pillared spire. Tube feet supporting tables with a short to medium-size spire................................... 4
3 Mean length of body wall tables 50 to 99 μm; width of spire at base 25 to 49 μm. Temperate distribution (
California
,
USA
)....................................................................
Pentamera charlottae
Deichmann, 1938
- Mean length of body wall tables less than 50 μm; width of spire at base less than 25 μm. Tropical distribution (
Costa Rica
).........................................................................
Pentamera beebei
Deichmann, 1938
4 Moderate calcareous ring, oval tables with four central holes in the body wall..................................... 5
- Long calcareous ring, triangular, rounded or acorn-shaped tables with ladder-shaped spire in the body wall.............. 6
5 Small oval buttons with two central and up to eight marginal knobs in body wall, tentacles with round to oblong plates with perforations and knobs..............................................
Pentamera montereyensis
Deichmann, 1938
- Body wall with thick oval, knobbed plates with meshwork of bumps covering one side, without buttons, tentacles with oblong reticulate plates......................................................
Pentamera trachyplaca
(H.L.
Clark, 1924
)
6 Body wall without triangular ossicles..................................................................... 7
- Body wall with triangular ossicles........................................................................ 8
7 Crowded layer of acorn-like cups with two-pillared, tapering spire rising from a cup-shaped base..............................................................................................
Pentamera zacae
Deichmann, 1938
- Without a crowded layer of acorn-like cups with two-pillared, tapering spire rising from a cup-shaped base.............. 9
8 Large, oval to triangular plates, rarely star-shaped tables, supporting tables with moderate spire, introvert with oval plates with serrated edge and blunt spines on surface..............................
Pentamera pseudocalcigera
Deichmann, 1938
- Circular to triangular or star-shaped tables with a wide central spire, supporting tables with low bumpy spire, introvert with elongated to oval plates with numerous bumps and raised central holes..................
Pentamera rigida
Lambert, 1998
9 Circular to oval tables, with smooth margin................................................................ 10
- Elongated tables, with wavy margin.........................................
Pentamera constricta
(
Ohshima, 1915
)
10 Introvert with plates and/or rosettes...................................................................... 11
- Introvert with tables......................................................
Pentamera pediparva
Lambert, 1998
11 Ossicles of body wall in one layer....................................................................... 12
- Ossicles of body wall in two layers.......................................
Pentamera lissoplaca
(H.L.
Clark, 1924
)
12 Tentacles with rosettes only................................................
Pentamera chiloensis
(
Ludwig, 1887
)
- Tentacles with plates only, or plates and rosettes............................................................ 13
13 Tentacles with elongated diamond-shaped plates with two large central holes, some with bumps or low pillar arch.............................................................................
Pentamera populifera
(
Stimpson, 1864
)
- Tentacles with plates and rosettes....................................................................... 14
14 Ossicles of body wall with tables with stout spire (diameter of the disk 70–100 μm), ending in a few teeth.............................................................................
Pentamera pseudopopulifera
Deichmann, 1938
- Ossicles of body wall with tables with ladder-shaped spire (diameter of the disk 30–50 μm)......
Pentamera fonsecae
n. sp.