New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae) Author Perkins, Philip D. text Zootaxa 2011 2011-10-28 3074 1 198 journal article 31530 10.5281/zenodo.1050060 2d28bebf-e3c5-457b-9152-8a6877ded281 1175-5326 1050060 Umbolenta Complex Three species comprise the Umbolenta Complex: H. umbolenta , H. kellymilleri , and H. loripes ( Figs. 108–110 ). These are small species (ca. 1.23–1.32 mm ), which have a pronotum with a distinct dark fascia, coarse dorsal punctation, lack a pronotal scintilla, and have very small plaques. The dorsal habitus is somewhat similar to that of H. concepcionica ( Fig. 119 ) in the Steineri Complex, but the pronotum has a fascia, the metaventral plaques are smaller, and the male genitalia are very different in basic form. The male genitalia in this complex are very distinctive, short and wide, with the parameres originating on the ventral face of the main piece, the left paramere much longer than the right, and the gonopore is located at the end of a long tube-shaped process that extends well beyond the other parts of the distal piece ( Figs. 111–113 ). The currently known species are from Paraguay ( H. umbolenta ), and Venezuela ( H. kellymilleri , and H. loripes ).