Order Chiroptera - Family Vespertilionidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 451 529 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Nyctalus leisleri subsp. verrucosus Bowditch 1825 Synonyms: Nyctalus leisleri subsp. madeirae Barrett-Hamilton 1906 .