Naticid gastropods from the middle Miocene of western Ukraine Author Pedriali, Luca Author Sosso, Maurizio Author Dell’Angelo, Bruno text Zootaxa 2019 2019-11-19 4700 2 151 195 journal article 24866 10.11646/zootaxa.4700.2.1 1a395b3c-4d5d-4b3b-b6e1-2e03784b3fcc 1175-5326 3548348 9B9BAB27-BFA6-4C74-809E-123C732A8890 Cochlis ukrainensis sp. nov. (Fig. 24.1−6) Type material. Holotype and 6 paratypes ( Table 5 ). Type locality. Varovtsi , Ukraine . Type stage. Middle Miocene ( Upper Badenian ) . Etymology. The name refers to Ukraine , the country where the village of Varovtsi is located. TABLE 5. Cochlis ukrainensis sp. nov. , type material, repository numbers and basic parameters. Spm stands for number of specimens; H stands for shell height (indicated as the maximum value if the paratype includes more than one shell)

type material	repository	locality	spm	material	H (mm)	figures
Holotype	MPUM 11735	Varovtsi	1	shell	35.41	24.1a–c
Paratype 1	MPUM 11736	Varovtsi	1	shell	21.27	24.2a–b
Paratype 2	MZB 32145	Varovtsi	1	shell	17.90	24.3
Paratype 3	ZISP 1/62193	Varovtsi	1	shell	26.27	24.4a–b
Paratype 4	IGS NANU B-III-3/2018	Varovtsi	1	shell	6.90	24.5
Paratype 5	MPUM 11737	Varovtsi	1	shell	21.22
Paratype 6	IGS NANU B-III-4/2018	Varovtsi	1	shell	17.56

Other material. Varovtsi: NP 10023 (2 shells); MZB 50516 (1 shell, Fig. 24.6); BD 164 (2 shells); MS 103 (2 shells). Horodok: NP 10022 (1 shell). TABLE 6. Cochlis ukrainensis sp. nov. , shell dimensions of the species subject to statistic analysis (10 specimens measured from Varovtsi). Notes: * = 2 protoconchs measured.

parameter	95° confidence interval	mean
DHW	0.189/0.200 mm *	0.194 mm
PD	0.767/0.789 mm *	0.778 mm
H	6.744−34.408 mm	20.576 mm
D	6.121−36.429 mm	21.275 mm
SH	1.183−8.811 mm	4.997 mm
AH	5.087−26.071 mm	15.579 mm
AW	4.393−20.965 mm	12.679 mm
UW	3.005−12.301 mm	7.653 mm
WUC	0.865−2.753 mm	1.809 mm
WAD	0.632−5.884 mm	3.258 mm
WAB	1.036−4.136 mm	2.586 mm
IS	10−30º	20º
SA	116−136º	126º

Diagnosis. Shell globose with depressed, low-conical spire and globular last whorl. Umbilicus deep, mediumsized. Parietal callus subrectangular, thick; anterior lobe flat and rounded. Funicle cord-like, thick; umbilical callus small, thick, semicircular in outline and separated from parietal callus by broad, regularly arched notch. Colour pattern of whitish to brown, uneven collabral stripes over a uniform light brown to brown background. FIGURE 24. Cochlis spp. from the Miocene of Ukraine and Italy, and from the Pliocene of Italy. 1–6. Cochlis ukrainensis sp. nov. from the middle Miocene (Badenian) of Ukraine at Varovtsi. 1. Holotype, MPUM 11735; 2. Paratype 1, MPUM 11736; 3. Paratype 2, MZB 32145; 4. Paratype 3, ZISP 1/62193; 5. Paratype 4, IGS NANU B-III-3/2018, protoconch; 6. MZB 50516, protoconch. 7–9. Cochlis raropunctata raropunctata ( Sasso, 1827 ) from Italy. 7. NP 10050, from the Pliocene of Pradalbino II, Bologna, with operculum in situ , and operculum by itself; 8. NP 10051 from the Pliocene of Balzo del Musico, Bologna, with operculum in situ , and operculum by itself; 9. MPUM 11245 from the Miocene (Tortonian) of Rio di Bocca d’Asino, Alessandria, protoconch (specimen figured by Robba et al . 2016 , pl. 3, fig. 3). Arrows point to the protoconch-teleoconch transition. Scale bars: 10 mm (figs 1–4, 7a, 8a, 8b); 5 mm (figs 7b, 8c); 500 µm (figs 5, 6, 9). Description. Protoconch small, depressed turbiniform, of 1.75 gently convex, apparently smooth whorls, tip medium-sized. Teleoconch globose, solid. Spire low-conical, rather depressed, whorls convex, suture slightly incised. Last whorl globular, moderately produced toward aperture, slightly expanded in a few young specimens. Subsutural shelf wide, flat or gently sloping, periphery above midline. Aperture D-shaped, in prosocline plane. Pari- etal callus moderately thick, subrectangular, with rounded, flat anterior lobe at level of umbilical border. Umbilicus deep, medium-sized. Funicle cord-like, moderately thick, separated from umbilical wall by narrow spiral furrow. Umbilical callus small, thick, semicircular in outline, located at abapical one-third of inner lip, separated from parietal callus by broad, regularly arched notch. Basal fasciole wide, depressed. Surface with rather dense, thin growth lines, stronger on subsutural shelf and basal fasciole; faint spiral striation present on body whorl. Background uniform light brown to brown, with pattern of uneven whitish collabral stripes. No opercula found. Remarks. Cochlis ukrainensis sp. nov. is quite similar to the late Miocene to early Pleistocene South European subspecies C. raropunctata raropunctata (Fig. 24.7−9), but differs from it in having: 1) a protoconch with 0.5 more whorls, with smaller diameter of the first half-whorl (average values 0.195 mm and 0.357 mm , respectively); 2) a wider, subrectangular parietal callus, with a better developed anterior lobe; 3) a smaller umbilicus; 4) an umbilical callus demarcated from the parietal callus by a broad, arched notch (reverse J-shaped in C. raropunctata raropunctata ); 5) a depressed basal fasciole, not sharply edged abaxially by a low step; and 6) a different and less variable colour pattern of uneven whitish to brown collabral stripes (a reddish-brown pattern of dots or spots, often arranged in collabral rows, spirally elongated spots, chevron marks, and irregular spiral broken lines in C. raropunctata raropunctata ). It is also worth noting that C. ukrainensis sp. nov. attains a smaller size. Distribution. Middle Miocene: Central Paratethys (Upper Badenian) in Ukraine (this paper).