Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)
Author
Oláh, J.
Author
Johanson, K. A.
text
Zootaxa
2008
2008-06-16
1802
1
248
journal article
11755334
Calosopsyche domingensis
(Banks)
Hydropsyche domingensis
Banks, 1941: 398
.
Plectropsyche domingensis
(
Banks, 1941
)
;
Ross & Unzicker 1977: 308
.
Hydropsyche domingensis
Banks, 1941
;
Botosaneanu 1996: 17
.
Calosopsyche domingensis
(
Banks, 1941
)
;
Flint, Holzenthal & Harris 1999: 75
.
Type
locality.
Santo Domingo
.
Remarks: As is seen from the frequent generic transfers, the status of this species was highly instable. Even in the cladistic analysis of
Schefter (2005)
it was suggested and isolated into a distinct genus. However, based upon defined primary generic characters, the species represents a true
Calosopsyche
, having identical primary generic characters with the other members of the genus. We have found some mistakes in the character matrix used in the cladistic analysis by
Schefter (2005)
. These mistakes were mainly caused by sexual dimorphism. The pretarsal claw is asymmetrical, not symmetrical. Like in other genera, or other species, even inside the genus
Calosopsyche
, asymmetry of the pretarsal claws is frequently sex linked. Even the degree of asymmetry varies greatly. Asymmetry involving laterally flanked setal bundle may occur only in the fore-leg, fore- and mid-leg, or in all legs in the male. This may, or may not be combined with identical variation in the female. The hind wing crossvein m-cu, representing one of the most important primary generic characters, is well visible in both
Calosopsyche continentalis
Flint & Bueno-Soria
, and
Calosopsyche domingensis
(Banks)
, like in all other species in the genus. Among the tertiary generic characters, the
endophallus
is long, not short as listed in the character matrix. Even the 3 setose processes are present, but shortened, forming almost a single, united, lobe. A careful examination shows that 3 separate setose surfaces are distinguishable, among them, the preanal appendages. After correcting the character matrix of
Schefter (2005)
for these characters, one of us (
KAJ
) re-ran the analysis in
PAUP
*. The many more shortest trees obtained were 2 steps shorter and the strict consensus tree did not improve resolution. Furthermore, the two
Calaosopsyche
species were still distantly related.