<document id="28ED4DA71362CEF9508FF57CCB060BF3" ID-DOI="10.11646/zootaxa.4098.3.1" ID-GBIF-Dataset="8770e7d9-65f4-4cee-a935-a5f365a95f32" ID-ISSN="1175-5326" ID-Zenodo-Dep="258895" ID-ZooBank="00F1BDB5-AB25-47A0-B789-2E05D2E683DE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460009663303" checkinUser="plazi" docAuthor="Beaver, Roger A., Ghahari, Hassan &amp; Sanguansub, Sunisa" docDate="2016" docId="6B5C9A7C475CFFD9C797E028FC07FF06" docLanguage="en" docName="zootaxa.4098.3.1.pdf.imf" docOrigin="Zootaxa 4098 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pityogenes quadridens Hartig 1834" docType="treatment" docVersion="10" lastPageNumber="414" masterDocId="9765E2044751FFD7C700E71CFFD5FFD3" masterDocTitle="An annotated checklist of Platypodinae and Scolytinae (Coleoptera: Curculionidae) from Iran" masterLastPageNumber="441" masterPageNumber="401" pageNumber="414" updateTime="1698233404894" updateUser="plazi">
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<mods:title id="FC4E68ABFC5F50396C1AAF10F3BF2B3C">An annotated checklist of Platypodinae and Scolytinae (Coleoptera: Curculionidae) from Iran</mods:title>
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<mods:namePart id="A06B146575752504A6C12C6F177F3449">Beaver, Roger A.</mods:namePart>
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<mods:namePart id="337FAA7C1162DD527B61C4377A09B976">Ghahari, Hassan</mods:namePart>
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<mods:namePart id="D9EC82AA6CF24F7271938BF8F25576C7">Sanguansub, Sunisa</mods:namePart>
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<mods:date id="AC1DCE66D7383CD0F27FC8B7828CEBFA">2016</mods:date>
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<treatment id="6B5C9A7C475CFFD9C797E028FC07FF06" ID-DOI="http://doi.org/10.5281/zenodo.5669446" ID-GBIF-Taxon="119381137" ID-Zenodo-Dep="5669446" LSID="urn:lsid:plazi:treatment:6B5C9A7C475CFFD9C797E028FC07FF06" httpUri="http://treatment.plazi.org/id/6B5C9A7C475CFFD9C797E028FC07FF06" lastPageId="14" lastPageNumber="414" pageId="13" pageNumber="414">
<subSubSection id="ABEF78E1475CFFDAC797E028FD9FF89E" box="[151,586,1844,1869]" pageId="13" pageNumber="414" type="nomenclature">
<paragraph id="E34A2B6A475CFFDAC797E028FD9FF89E" blockId="13.[151,586,1844,1869]" box="[151,586,1844,1869]" pageId="13" pageNumber="414">
<heading id="B8029C06475CFFDAC797E028FD9FF89E" bold="true" box="[151,586,1844,1869]" fontSize="10" level="2" pageId="13" pageNumber="414" reason="2">
<taxonomicName id="24F550E9475CFFDAC797E028FD9FF89E" ID-CoL="4JCLQ" authority="Hartig, 1834" authorityName="Hartig" authorityYear="1834" box="[151,586,1844,1869]" class="Insecta" family="Curculionidae" genus="Pityogenes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="D181F778475CFFDAC797E028FD9FF89E" bold="true" box="[151,586,1844,1869]" pageId="13" pageNumber="414">
<emphasis id="D181F778475CFFDAC797E028FE4EF89E" bold="true" box="[151,411,1844,1869]" italics="true" pageId="13" pageNumber="414">Pityogenes quadridens</emphasis>
(Hartig, 1834)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ABEF78E1475CFFD9C797E060FC07FF06" lastPageId="14" lastPageNumber="415" pageId="13" pageNumber="414" type="distribution">
<paragraph id="E34A2B6A475CFFDAC797E060FCD4F847" blockId="13.[151,1436,1916,2013]" box="[151,769,1916,1941]" pageId="13" pageNumber="414">
<emphasis id="D181F778475CFFDAC797E060FE5FF846" bold="true" box="[151,394,1916,1941]" pageId="13" pageNumber="414">
Distribution in 
<collectingCountry id="9BE26BFA475CFFDAC64EE060FE50F846" box="[334,389,1916,1941]" name="Iran" pageId="13" pageNumber="414">Iran</collectingCountry>
.
</emphasis>
Mazandaran (
<bibRefCitation id="8764569B475CFFDAC52AE060FD20F847" author="Samin" box="[554,757,1916,1941]" pageId="13" pageNumber="414" refString="Samin, N., Sakenin, H. &amp; Rastegar, J. (2011) A study of the species composition of Scolytidae (Coleoptera) of north and northwestern Iran. Amurian Zoological Journal, 3, 265 - 267." type="journal article" year="2011">
Samin 
<emphasis id="D181F778475CFFDAC57BE061FD79F846" box="[635,684,1916,1941]" italics="true" pageId="13" pageNumber="414">et al</emphasis>
. 2011
</bibRefCitation>
).
</paragraph>
<paragraph id="E34A2B6A475CFFD9C7C7E0BCFC07FF06" blockId="13.[151,1436,1916,2013]" lastBlockId="14.[151,1436,151,213]" lastPageId="14" lastPageNumber="415" pageId="13" pageNumber="414">
<emphasis id="D181F778475CFFDAC7C7E0BCFE16F86A" bold="true" box="[199,451,1952,1977]" pageId="13" pageNumber="414">General distribution.</emphasis>
Europe, through 
<collectingCountry id="9BE26BFA475CFFDAC586E0BCFD04F86A" box="[646,721,1952,1977]" name="Russia" pageId="13" pageNumber="414">Russia</collectingCountry>
to the Far East, 
<collectingCountry id="9BE26BFA475CFFDAC489E0BCFC0FF86A" box="[905,986,1952,1977]" name="Turkey" pageId="13" pageNumber="414">Turkey</collectingCountry>
, 
<collectingCountry id="9BE26BFA475CFFDAC4E4E0BCFBBEF86A" box="[996,1131,1952,1977]" name="Kazakhstan" pageId="13" pageNumber="414">Kazakhstan</collectingCountry>
. 
<emphasis id="D181F778475CFFDAC7C7E0D8FEF2F80E" bold="true" box="[199,295,1988,2013]" pageId="13" pageNumber="414">Biology.</emphasis>
The species usually breeds in species of 
<taxonomicName id="24F550E9475CFFDAC41AE0D9FC8FF80F" box="[794,858,1989,2012]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="13" pageNumber="414" phylum="Tracheophyta" rank="genus">
<emphasis id="D181F778475CFFDAC41AE0D9FC8FF80F" box="[794,858,1989,2012]" italics="true" pageId="13" pageNumber="414">Pinus</emphasis>
</taxonomicName>
, but occasionally attacks 
<taxonomicName id="24F550E9475CFFDAC38EE0D8FB19F80E" box="[1166,1228,1988,2013]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="13" pageNumber="414" phylum="Tracheophyta" rank="genus">
<emphasis id="D181F778475CFFDAC38EE0D8FB19F80E" box="[1166,1228,1988,2013]" italics="true" pageId="13" pageNumber="414">Abies</emphasis>
</taxonomicName>
, 
<taxonomicName id="24F550E9475CFFDAC3DFE0D9FAC9F80F" box="[1247,1308,1989,2012]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="13" pageNumber="414" phylum="Tracheophyta" rank="genus">
<emphasis id="D181F778475CFFDAC3DFE0D9FAC9F80F" box="[1247,1308,1989,2012]" italics="true" pageId="13" pageNumber="414">Larix</emphasis>
</taxonomicName>
and 
<taxonomicName id="24F550E9475CFFD9C25CE0D9FED8FF63" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" lastPageId="14" lastPageNumber="415" order="Pinales" pageId="13" pageNumber="414" phylum="Tracheophyta" rank="genus">
<emphasis id="D181F778475CFFDAC25CE0D9FA49F80F" box="[1372,1436,1989,2012]" italics="true" pageId="13" pageNumber="414">Picea</emphasis>
(Pinaceae)
</taxonomicName>
(
<bibRefCitation id="8764569B475FFFD9C61BE78BFE4CFF63" author="Stark" box="[283,409,151,176]" pageId="14" pageNumber="415" refString="Stark, V. N. (1952) Fauna of the USSR, Coleoptera 31, Bark Beetles. Akademia Nauk, Moscow, 463 pp. [in Russian]" type="book" year="1952">Stark 1952</bibRefCitation>
, 
<bibRefCitation id="8764569B475FFFD9C6A4E78BFDE1FF63" author="Schedl" box="[420,564,151,176]" pageId="14" pageNumber="415" refString="Schedl, K. E. (1962) Bestimmungstabellen palaearktischer Borkenkafer IX. Centralblatt fur das Gesamte Forstwesen, 79, 132 - 159." type="journal article" year="1962">Schedl 1962</bibRefCitation>
). Recorded in 
<collectingCountry id="9BE26BFA475FFFD9C5D5E784FCD7FF7C" box="[725,770,152,175]" name="Iran" pageId="14" pageNumber="415">Iran</collectingCountry>
from 
<taxonomicName id="24F550E9475FFFD9C447E785FBA3FF63" box="[839,1142,151,176]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="14" pageNumber="415" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="D181F778475FFFD9C447E785FC2DFF63" box="[839,1016,151,176]" italics="true" pageId="14" pageNumber="415">Picea orientalis</emphasis>
(Pinaceae)
</taxonomicName>
(
<bibRefCitation id="8764569B475FFFD9C384E78BFA9AFF63" author="Samin" box="[1156,1359,151,176]" pageId="14" pageNumber="415" refString="Samin, N., Sakenin, H. &amp; Rastegar, J. (2011) A study of the species composition of Scolytidae (Coleoptera) of north and northwestern Iran. Amurian Zoological Journal, 3, 265 - 267." type="journal article" year="2011">
Samin 
<emphasis id="D181F778475FFFD9C3D4E785FAD0FF63" box="[1236,1285,151,176]" italics="true" pageId="14" pageNumber="415">et al</emphasis>
. 2011
</bibRefCitation>
). 
<bibRefCitation id="8764569B475FFFD9C261E78BFF34FF06" author="Stark" pageId="14" pageNumber="415" refString="Stark, V. N. (1952) Fauna of the USSR, Coleoptera 31, Bark Beetles. Akademia Nauk, Moscow, 463 pp. [in Russian]" type="book" year="1952">Stark (1952)</bibRefCitation>
and 
<bibRefCitation id="8764569B475FFFD9C618E7A0FE63FF06" author="Schedl" box="[280,438,188,213]" pageId="14" pageNumber="415" refString="Schedl, K. E. (1962) Bestimmungstabellen palaearktischer Borkenkafer IX. Centralblatt fur das Gesamte Forstwesen, 79, 132 - 159." type="journal article" year="1962">Schedl (1962)</bibRefCitation>
briefly describe the biology and gallery system.
</paragraph>
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