Review of the Blastobasinae of Costa Rica (Lepidoptera: Gelechioidea: Blastobasidae) Author Adamski, David text Zootaxa 2013 3618 1 1 223 journal article 39040 10.11646/zootaxa.3618.1.1 d2a7d193-4512-4565-a56e-49034dcf6101 1175-5326 247396 B548B139-E8D9-4F10-956E-E0001E6C7586 Key to the Species of Hypatopa of Costa Rica 1. Male................................................................................................ 2 - Female............................................................................................. 50 2. Gnathos present)...................................................................................... 3 - Gnathos absent).......................................................................... Hypatopa rabio 3. Gnathos emarginate mesially ( Figs. 139 , 145 , 149, 151 , 153, 157 , 159, 161, 163 , 165, 167, 169 , 171 , 177, 179, 181 , 183, 185 , 189, 191 , 195 , 201, 203, 205 , 209, 211 , 213, 215 , 219, 221, 223 , 225 , 231 )........................................ 4 - Gnathos not emarginate mesially ( Figs. 137 , 147 , 173 , 141, 143 , 155 , 175 , 187 , 193 , 197, 199 , 207 , 217 , 227 , 231, 233 )... 36 4. Mesial emargination on a slightly raised or protuberant lobe ( Figs. 139 , 145 , 153 , 163 , 171 , 183 , 189 , 203, 205 , 209 , 221 , 225 , 231 )................................................................................................ 5 - Mesial emargination on an entire margin ( Figs. 149, 151 , 157 , 159, 161 , 167, 169 , 177, 179, 181 , 185 )................ 17 5. Apical process of ventral part of valva acutely curved in area from base to 1/2 ( Figs. 139 , 163 , 209 , 231 )................ 6 - Apical process of ventral part of valva not acutely curved in area from base to 1/2 ( Figs. 145 , 153 , 171 , 183 , 189 , 203, 205 , 221 , 225 )............................................................................................ 9 6. Apical process of ventral part of valva acutely curved laterally ( Figs. 231 )............................. Hypatopa rudis - Apical process of ventral part of valva not acutely curved laterally (139, 163, 209).................................. 7 7. Proximal flange of valva longer than wide ( Fig. 163 ); anellus rounded apically ( Fig. 164 )............... Hypatopa plebis - Proximal flange of valva not longer than wide ( Figs. 139 , 209 ); anellus emarginate apically ( Figs. 140 , 210 )............. 8 8. Gnathos deeply emarginate mesially; proximal flange wider than long ( Fig. 139 )...................... Hypatopa cladis - Gnathos shallowly emarginate mesially; proximal flange about as wide as long ( Fig. 209 ).................. Hypatopa io 9. Margin of proximal flange of valva crenulate ( Figs. 183 , 225 )................................................. 10 - Margin of proximal flange of valva serrate ( Figs. 145 , 153 , 171 , 189 , 203, 205 , 221 )............................... 11 10. Gnathos deeply emarginate mesially ( Fig. 183 ); anellus truncate apically ( Fig. 184 )................... Hypatopa semela - Gnathos shallowly emarginate mesially ( Fig. 225 ); anellus deeply emarginate, forming two lateral lobes and with two porrect median spinelike processes ( Fig. 226 )......................................................... Hypatopa solea 11. Gnathos deeply emarginate mesially ( Figs. 153 , 171 )........................................................ 12 - Gnathos shallowly emarginate mesially ( Figs. 145 , 189 , 203, 205 , 221 ).......................................... 13 12. Sclerite of phallus acutely curved near middle; vesica with barbed cornuti ( Fig. 154 )..................... Hypatopa crux - Sclerite of phallus shallowly curved near middle; vesica without cornuti ( Fig. 172 ).................... Hypatopa caepae 13. Phallus and sclerite of phallus longer than valva ( Figs. 145 , 204, 206 )........................................... 14 - Phallus and sclerite of phallus shorter than valva ( Figs. 190 , 222 )............................................... 16 14. Sclerite of phallus acutely sigmoid-shaped ( Fig. 204 )............................................... Hypatopa dux - Sclerite of phallus not acutely curved throughout length ( Figs. 146 , 206 )......................................... 15 15. Juxta divided mesially ( Fig. 145 ); anellus nearly parallelsided throughout most of length ( Fig. 146 )........ Hypatopa cotis - Juxta bandlike ( Fig. 205 ); anellus constricted beyond base ( Fig. 206 )................................. Hypatopa erato 16. Gnathos widely emarginate ( Fig. 189 ); anellus near parallelsided throughout most of length ( Fig. 190 )...... Hypatopa rego - Gnathos narrowly emarginate ( Fig. 221 ); anellus gradually narrowed from base ( Fig. 222 )............... Hypatopa umbra 17. Gnathos projected anteriorly near or greater than 1/2 width ( Figs. 169 , 177 , 191 , 195 , 201 , 219, 223 ).................. 18 - Gnathos not projected anteriorly as above ( Figs. 149, 151 , 157 , 159, 161 , 165, 167 , 179, 181 , 185 , 211 , 213, 215 )........ 24 18. Ventral part of valva greatly protracted inwards basally ( Figs. 169 , 195 , 23 )...................................... 19 - Ventral part of valva slightly protracted inwards basally ( Figs. 177 , 191 , 201 , 19 ).................................. 21 19. Proximal flange of valva nearly twice as long as wide; apical process of ventral part of valva acutely curved basally ( Fig. 169 )....................................................................................... Hypatopa manus - Proximal flange of valva about as long as wide; apical process of ventral part of valva not acutely curved from base ( Figs. 195 , 223 )............................................................................................... 20 20. Apical margin of proximal flange wider than base ( Fig. 223 ); anellus widely emarginate, forming two lateral lobes, and with two convergent apical processes ( Fig. 224 )...................................................... Hypatopa texo - Apical margin of proximal flange narrower than base ( Fig. 195 ); anellus rounded apically ( Fig. 196 )........ Hypatopa verax 21. Gnathos deeply emarginate mesially ( Figs. 177 , 191 , 219 ).................................................... 22 - Gnathos shallowly emarginate mesially ( Fig. 201 ).................................................. Hypatopa nox 22. Sclerite of phallus recurved apically ( Figs. 192 , 220 )......................................................... 23 - Sclerite of phallus not recurved apically ( Fig. 178 ).............................................. Hypatopa scobis 23. Margin of proximal flange serrate ( Fig. 191 ); sclerite of phallus recurved to near base of anellus ( Fig. 220 ).... Hypatopa ira - Margin of proximal flange entire ( Fig. 219 ); sclerite of phallus recurved to subapical region ( Fig. 192 )...... Hypatopa styga 24. Ventrolateral margin of proximal flange with two subequal, large spinelike processes ( Fig. 149 ).......... Hypatopa hecate - Ventrolateral margin of proximal flange entire, crenulate, or serrate ( Figs. 151 , 157 , 159, 161 , 165, 167 , 179, 181 , 185 , 211 , 213, 215 )........................................................................................... 25 25. Margin of proximal flange entire or crenulate.............................................................. 26 - Margin of proximal flange serrate........................................................................ 30 26. Base of phallus widely bulbous basally ( Figs. 152 , 180 , 214 ).................................................. 27 – Base of phallus not widely bulbous basally ( Figs. 160 , 166 )................................................... 29 27. Vesica with cornuti ( Fig. 152 )................................................................. Hypatopa acus - Vesica without cornuti ( Fig. 180 , 214 ).................................................................... 28 28. Proximal flange elongate ( Fig. 213 ); sclerite of phallus straight basally ( Fig. 214 )......................... Hypatopa vox - Proximal flange ellipsoid ( Fig. 179 ); sclerite of phallus acutely curved basally ( Fig. 180 )................. Hypatopa agnae 29. Anellus 1/2 length of phallus, nearly parallelsided ( Fig. 166 )....................................... Hypatopa dolo - Anellus less than 1/2 length of phallus, wider on basal 2/3, abruptly narrowed apically ( Fig. 160 )......... Hypatopa arxcis 30. Juxta divided mesially ( Figs. 161 , 211 ).................................................................... 31 - Juxta bandlike ( Figs. 157 , 167 , 181 , 185 , 215 ).............................................................. 32 31. Apical process of ventral part of valva downcurved ( Fig. 161 ); anellus broadly rounded apically ( Fig. 162 ).... Hypatopa caedis - Apical process of ventral part of valva upturned ( Fig. 211 ); anellus broadly emarginate apically ( Fig. 212 ).... Hypatopa leda 32. Anellus apically emarginate ( Figs. 158 , 182 , 216 )........................................................... 33 - Anellus apically rounded ( Figs. 168 , 186 )................................................................. 35 33. Anellus shallowly emarginate apically, bulbous basally ( Fig. 158 )................................... Hypatopa hera - Anellus deeply emarginate apically, not bulbous basally ( Figs. 182 , 216 )......................................... 34 34. Emargination of anellus forming two apically rounded lateral lobes ( Fig. 182 )........................ Hypatopa phoebe - Emargination of anellus forming two convergent, apically pointed lateral lobes ( Fig. 216 )............ Hypatopa tapadulcea 35. Proximal flange subrectangular; spinelike process of ventral part of valva rounded apically ( Fig. 167 )....... Hypatopa cyane - Proximal flange semicircular; apical process of ventral part of valva acute ( Fig. 185 )..................... Hypatopa edax 36. Gnathos with a small or protuberant mesial lobe ( Figs. 137 , 147 , 155 , 173 , 187 )................................... 37 - Gnathos with mesial lobe absent ( Figs. 141, 143 , 175 , 193 , 197, 199 , 207 , 217 , 227 , 233 )........................... 41 37. Proximal flange of valva with 1–4 spinelike processes on apicoventral region ( Figs. 148 , 174 )........................ 38 - Proximal flange of valva without such processes ( Figs. 137 , 155 , 187 )........................................... 39 38. Proximal flange of valva elongate, densely setose ( Fig. 173 ); sclerite of phallus broadly curved ( Fig. 174 ).. Hypatopa cotytto - Proximal flange of valva not elongate, sparsely setose ( Fig. 147 ); sclerite of phallus acutely curved at 2/3 ( Fig. 148 )................................................................................................. Hypatopa pica 39. Margin of proximal flange of valva entire ( Fig. 137 ); anellus with apical barb ( Fig. 138 )................... Hypatopa nex - Margin of proximal flange serrate ( Figs. 155 , 187 ); anellus without apical barb ( Figs. 156 , 188 )...................... 40 40. Proximal flange serrate ( Fig. 155 ); apical margin of anellus narrower than base ( Fig. 156 )................ Hypatopa limae - Proximal flange deeply serrate ( Fig. 187 ); anellus truncate apically ( Fig. 188 )........................ Hypatopa joniella 41. Apical process of ventral part of valva diminutive ( Figs. 141, 143 ); apex of anellus bearing a pair of recurrent lateral barblike processes ( Figs. 142, 144 ).............................................................................. 42 - Apical process of ventral part of valva of normal length ( Figs. 197, 199 , 207 , 217 , 227 , 233 ); anellus not bearing recurrent barblike processes ( Figs. 198, 200 , 208 , 218 , 228 , 234 )....................................................... 43 42. Anellus with crenulate barblike processes ( Fig. 142 ).............................................. Hypatopa juno - Anellus with straight barblike processes ( Fig. 144 )............................................... Hypatopa actes 43. Anellus mesially emarginate apically ( Figs. 228 , 234 )........................................................ 44 - Anellus not mesially emarginate apically ( Figs. 176 , 194 , 198, 200 , 208 , 218 )..................................... 45 44. Anellus marginate, forming two large, divergent apicolateral processes ( Fig. 228 )................... Hypatopa bilobata - Anellus marginate, forming two short apicolateral processes ( Fig. 234 )............................... Hypatopa musa 45. Phallus widely bulbous basally ( Figs. 194 , 198, 200 )......................................................... 46 - Phallus not widely bulbous ( Figs. 176 , 208 , 218 )............................................................ 48 46. Apical part of sclerite of phallus slightly curved ( Fig. 194 ).......................................... Hypatopa texla - Apical part of sclerite of phallus recurved ( Figs. 198, 200 ).................................................... 47 47. Basal ridge of digitate process of dorsal part of valva densely setose; base of apical process of ventral part of valva with a set- ose lobe ( Fig. 199 )......................................................................... Hypatopa mora - Basal ridge of digitate process of dorsal part of valva sparsely setose; base of apical process of ventral part of valva without a setose lobe ( Fig. 197 ).................................................................. Hypatopa tapadulcea 48. Apical process of ventral part of valva downcurved ( Fig. 217 )...................................... Hypatopa dicax - Apical process of ventral part of valva not downcurved ( Figs. 175 , 207 )......................................... 49 49. Margin of proximal flange entire ( Fig. 175 ); sclerite of phallus shorter than phallus ( Fig. 176 )............ Hypatopa lucina - Margin of proximal flange serrate ( Fig. 207 ); sclerite of phallus as long as phallus ( Fig. 208 )................ Hypatopa fio 50. Eighth tergum with a darkly pigmented median longitudinal streak ( Figs. 265 , 270 , 272–273 )........................ 51 - Eighth tergum without a darkly pigmented median longitudinal streak ( Figs. 261–264 , 266–269, 271 ).................. 54 51. Ductus bursae coiled throughout most of length ( Fig.265 )...................................... Hypatopa tapadulcea - Ductus bursae not coiled throughout most of length ( Figs. 270 , 272–273 )........................................ 52 52. Posterior margin of seventh sternum emarginate ( Figs. 270 , 272 )............................................... 53 - Posterior margin of seventh sternum straight ( Fig. 273 )............................................. Hypatopa vitis 53. Posterior margin of seventh sternum laterally emarginate, forming a broad, truncate, median lobe ( Fig. 270 ).... Hypatopa rea - Posterior margin of seventh sternum mesially and laterally emarginate, forming two rounded lobes ( Fig. 272 )... Hypatopa gena 54. Sex scales on membrane adjacent to Ostium bursae or on margin of seventh sternum ( Figs. 261–262 , 268 ).............. 55 - Sex scales absent ( Figs. 263–264 , 266–267 , 269, 271 )........................................................ 57 55. Sex scales on membrane adjacent to Ostium bursae ( Fig. 261 )..................................... Hypatopa umbra - Sex scales on or near posterior margin of seventh sternum only ( Figs. 262 , 268 )................................... 56 56. Sex scales on posterolateral margin of seventh sternum; antrum present ( Fig. 262 )..................... Hypatopa hecate - Sex scales along entire posterior margin of seventh segment; antrum absent ( Fig. 268 )................... Hypatopa rabio 57. Signum present ( Figs. 264 , 266 , 269, 271 )................................................................. 59 - Signum absent ( Fig. 263 , 267 ).......................................................................... 58 58. Eighth sternum subquadrate; ductus seminalis anterior to anterior margin of seventh sternum ( Fig. 267 )...... Hypatopa texo - Eighth sternum longer than wide; ductus seminalis posterior to anterior margin of seventh sternum ( Fig. 263 )..................................................................................................... Hypatopa plebis 59. Eighth sternum quadrate ( Fig. 269 )............................................................. Hypatopa sais - Eighth sternum longer than wide ( Figs. 264 , 266 , 271 )....................................................... 60 60. Antrum present ( Figs. 264 , 266 )......................................................................... 61 - Antrum absent ( Fig. 271 ).................................................................... Hypatopa hora 61. Antrum wider than long; anterior 1/4 of ductus bursae spinulate ( Fig. 266 )............................. Hypatopa leda - Antrum longer than wide; anterior 1/2 of ductus bursae with two rows of internal platelets ( Fig. 264 )...... Hypatopa phoebe