A Revision Of The Palaearctic Species Of The Eugraphe Hübner, [1821] 1816 Generic Complex. Part I. The Genera Eugraphe And Goniographa (Lepidoptera, Noctuidae)
Author
Varga, Z.
Author
Ronkay, L.
text
Acta Zoologica Academiae Scientiarum Hungaricae
2002
2003-02-28
48
4
333
374
journal article
10.5281/zenodo.12587286
2064-2474
12587286
Goniographa discussa
sp. n.
(
Figs 4
,
26, 27
,
53, 54
)
Holotype
: male,
Seravshan Mts
,
45 km
SEE Aini
,
2000–2600 m
,
68°03’E
,
39°20’N
,
17–18. VII.1994
, leg.
LUKHTANOV
; slide No.
RL
7544m
(coll. P.
GYULAI
, in
HNHM
Budapest
).
Paratypes
.
Tadjikistan
:
1 female
,
Pamir Mts
,
Artuch valley
,
2–10.VIII.1988
, coll. A.V.
NEKRASOV
(coll.
HNHM
Budapest);
Pamir Mts
, Shugnan,
VI.1935
, leg.
WEIDINGER
(coll. G.
RONKAY
); vic of
Shuroabad
,
35 km
S Kuliabad
,
2000 m
,
12.VIII.1958
, leg.
SHCHETKIN
;
Peter
1st Mts, Chasor Thasma,
2100 m
,
13.VII.1977
, leg.
SHCHETKIN
;
Peter
1st Mts, Muk,
2100 m
,
27.VIII.1975
, leg.
SHCHETKIN
;
1 male
,
2 females
Peter
1st
Mts
,
Daran-Nazarak valley
,
Ganishou
,
2110 m
,
19–21.VIII. 1977
, leg.
SHCHETKIN
;
18 specimens
, from the same locality, VII-VIII.1994, leg.
SHCHETKIN
;
1 m
,
Peter
1st
Mts
,
Ganishou
,
1500–2500 m
,
11.VII.2000
, leg.
RYBAK
(coll.
LEHMANN
)
;
3 females
,
Daran-Nazarak valley
,
1700 m
,
15.VIII.1992
, leg.
SHCHETKIN
;
1 male
,
3 females
,
Karategin
range,
Sangikar
gorge,
1700 m
,
28.VIII.1969
, leg.
SHCHETKIN
;
2 males
,
2 females
,
Varzob valley
,
Maihur
,
2000 m
,
24.VII.–16.IX.1967
, leg.
SHCHETKIN
;
1 female
,
Hissar Mts
,
Anzob
pass,
3400 m
,
50 km
N
Acta zool. hung. 48, 2002
Dushanbe
,
24–25.VII.1994
, leg.
LUKHTANOV
;
2 males
, from the same locality
,
26.VIII.1967
, leg.
SHCHETKIN
;
2 males
,
Hissar Mts
,
25 km
S Pendzhikent
,
1800 m
,
10.VII.1994
, leg.
LUKHTANOV
;
1 female
,
Hissar Mts
,
Gushary
,
1300 m
,
18–25.IX.1985
, leg.
SHCHETKIN
(coll. A.
BECHER
, J.
STUMPF
and P.
GYULAI
).
Afghanistan
:
1 male
,
Badakhshan
, “Sarakanda”
4100 m
,
01.18.1953
. leg KLAP-
PERICH
(
ZSM
)
.
Slide Nos RL
7563m
, RL
7564m
, RL
7566m
, VZ4673, VZ7022 (males), RL7481f, RL7570f, RL7575f (females).
Diagnosis.
G. discussa
has narrower and more rounded forewings comparing with that of
G. decussa
, with generally more obsolescent markings. The maculation of the new species is less conspicuous, with less sharply defined ochreous outlines of the reniform and orbicular stigmata; the claviform stigma is narrower, more obsolescent. The dark intermacular patch of the cell, the basal dash and the subterminal arrowheads are also less sharply marked; the darker suffusion of the hindwing is more diffuse.
Comparing the male genitalia of the two sister species, the uncus of
G. discussa
is slightly broader than in
G. decussa
, slightly spatulate terminally; the valvae have a less convex dorsal costa; the harpe is slightly more elongate; the dentate, ribbon-like extension of the carina is only slightly arcuate, recurved terminally; the bulbed cornutus is more acute and about twice as large as in
G. decussa
.
Fig. 4.
Goniographa discussa
sp. n.
, paratypes, slide Nos a–b = VZ4673, c = RL7566, d = RL7564
Acta zool. hung. 48, 2002
The female genitalia of
G. discussa
differ from those of
G. decussa
and
G. shchetkini
by their characteristic, long and deep, sometimes sinuously folded ventral crest of the anterior third of ductus bursae, terminated in a large, more or less globular ventral pouch, both related species have a deeper or shallower arch at anterior edge of ventral plate instead of this structure (see
Figs 24–29
). The ostium bursae is smaller than that of
G. decussa
but larger than in
G. shchetkini
, the medio-caudal incision is the smallest in
G. discussa
. The anterior third of the ductus bursae is less dilated, its lateral margins are almost parallel, this part is broader (
G. decussa
)or conspicuously broader (
G. shchetkini
)in the two related species.
Description. Wingspan
30–35 mm
, length of forewing
13–15 mm
. The main external features fit well with those of the
decussa
-group, the differential characteristics are given in the diagnosis. The
holotype
(male)and a typical female are illustrated in
Figs 53 and 54
.
Male genitalia (
Fig. 4
): Uncus slightly broader, slightly spatulate terminally; dorsal costa of valva only slightly convex; harpe small, more elongate; pollex long and curved; juxta shield-shaped with narrow and shallow incision apically; aedeagus straight, long and strong; dentate ribbon slightly arcuate, recurved terminally; bulbed cornutus large, acute.
Female genitalia (
Figs 26, 27
): Ovipositor medium-long, rather weak; gonapophyses slender, fine. Ostium bursae sclerotized, its ventral plate relatively short, broadly triangular, caudal edge almost straight, with minute medio-caudal incision only. Ductus bursae strongly, granulously sclerotized, more or less flattened, long, broad, with almost parallel lateral margins, only posterior third dilated slightly. Ventral surface of ductus bursae with fine longitudinal medio-lateral suture, anterior third with long and deep, sometimes sinuously folded large crest running into a rather globular, big ventral pouch. Apical part of bursa copulatrix small, flattened-conical, wrinkled, partly gelatinous and scobinate. Appendix bursae rather long, elliptical, finely wrinkled and gelatinous; corpus bursae elliptical-saccate; signa absent.
Bionomics and distribution.
G. discussa
is distributed from the Zeravshan and Hissar Mts through the western Pamir Mts to NE
Afghanistan
(Prov.
Badakhshan
), mostly at medium but exceptionally also at rather high elevations.
Etymology. The specific name refers to the less sharply defined markings of the new species.