The Nearctic Ecclisomyia species (Trichoptera: Limnephilidae) Author Givens, Donald R. text Zootaxa 2018 2018-04-23 4413 2 201 259 journal article 30205 10.11646/zootaxa.4413.2.1 fa65180b-5ae2-4eb7-8616-1c131cfe7251 1175-5326 1227343 495CE6EB-4A83-4A05-8E42-A92889F1C1C4 Keys to western North American species of Ecclisomyia Males 1. Inferior appendages each without single black basal spine or mesal stout spines; phallus with pair of distinct sclerotized ventral parameres hooked distally ( Figs 1a , 2a )...................................................... E. bilera Denning - Inferior appendages each with single black basal spine or stout mesal spines ( Figs. 10a, 10b , 11b , 22b , 23b , 30b , 31b ); phallus with distinct pair of sclerotized parameres rounded distally ( Fig. 11a ), or with fused parameres ( Figs. 23a , 31a )........... 2 2. Inferior appendages each with stout mesal spine ( Figs.10a , 11b ); phallus with pair of elongate parameres, rounded distally ( Figs. 11a )............................................................................ E. conspersa Banks - Inferior appendages each with 2 or more stout, mesal spines; phallus with shorter and fused parameres ( Figs. 22a, 22b , 23a, 23b , 30a, 30b , 31a, 31b )................................................................................ 3 3. Apex of fused parameres with shallow U-shaped mesal indentation; inferior appendages long, lateral sides nearly parallel in ventral view ( Figs. 22b , 23b )............................................................. E. maculosa Banks - Apex of fused parameres with deep U-shaped mesal indentation; inferior appendages short, lateral sides indented basally in ventral view ( Figs. 30b , 31b ).............................................................. E. simulata Banks Females 1. Distal margin of tergum X in ventral view with shallow mesal indentation ( Figs. 12 , 13a ); spermathecal sclerite narrow and long ( Fig. 13a )........................................................................ E. conspersa Banks - Distal margin of tergum X in ventral view with V-shaped incision ( Figs. 3c , 4a, 4c , 24c , 25b , 32a, 32b ); spermathecal sclerite short, rounded, or quadrate ( Figs. 4a , 25b, 25c , 32b ).......................................................... 2 2. External gonopod of segment VIII longer than wide, its distal margin with shallow concave V-shaped indentation; external gonopod of segment IX broadly concave, without posteriorly directed lobes ( Figs. 3b , 4a ); spermathecal sclerite rounded ( Fig. 3b , 4a )................................................................................ E. bilera Denning - External gonopod of segment VIII at least as wide as long, distal margin with V-shaped incision ( Figs. 25b ), convex ( Fig. 24c ), straight ( Fig. 25c ), or slightly concave ( Fig. 32a ); external gonopod of segment IX with deep V-shaped incision, with posteriorly directed lobes ( Figs. 24c , 25b, 25c , 32a, 32b ); spermathecal sclerite quadrate (Figs, 24c, 25c, 32a, 32b).............. 3 3. External gonopod of segment IX long, narrow, deeply cleft extending posterad about 2 X its width beyond caudal margin of external gonopod of segment VIII ( Figs. 24c , 25b, 25c ); distal margin of tergum X in ventral view with V-shaped incision for about 3/4 length of tergite ( Figs. 24c , 25b, 25c ).............................................. E. maculosa Banks - External gonopod of segment IX short, wide, with V-shaped incision, only extending posterad 1 X its width beyond caudal margin of external gonopod of segment VIII ( Figs. 32a, 32b ); distal margin of tergum X in ventral view with deep V-shaped incision for about 1/2 length of tergite ( Figs. 32a, 32b ).......................................... E. simulata Banks Pupae 1. Apical processes each straight, directed caudad, with 3 long, stout black setae at apex ( Figs. 15a, 15b , 16a, 16b, 17a, 17b); lat- eral line gills on segments IIp–IVa ( Fig. 42 )................................................. E. conspersa Banks - Apical processes of females curved mesad, of males sometimes straight ( Figs. 33a, 33b ); each with 2 long stout black setae at apex ( Figs. 5a, 5b , 6a, 6b , 7a , 7b , 26a, 26b 27a, 27b , 28a , 28b , 33a, 33b , 34a, 34b , 35a, 35b); without lateral line gills ( Fig. 42 ).................................................................................................... 2 2. Abdominal segment VII without dorsoanterior or ventroanterior gills ( Fig. 42 )...................... E. maculosa Banks - Abdominal segment VII with dorsoanterior and ventroanterior gills............................................. 3 3. Dorsal abdominal segment II with anterior gills ( Fig. 42 )........................................ E. simulata Banks - Dorsal abdominal segment II without anterior gills ( Fig. 42 )..................................... E. bilera Denning Larvae 1. Head, thoracic sclerites uniformly dark brown, without mesal yellow striping....................... E. bilera Denning - Head, thoracic sclerites with mesal yellowish striping........................................................ 2 2. With lateral line gills on segments IIp–IVa ( Fig. 42 )........................................... E. conspersa Banks - Without lateral line gills ( Fig. 42 )........................................................................ 3 3. Without anterior gills on dorsal and ventral abdominal segment VII; generally lacking dorsoanterior gills on abdominal seg- ment II ( Fig. 42 )....................................................................... E. maculosa Banks - With anterior gills on dorsal and ventral abdominal segment VII; with dorsoanterior gills on abdominal segment II ( Fig. 42 )....................................................................................... E. simulata Banks