Revision of the Hylicellidae of the Late Triassic of Queensland (Hemiptera Cicadomorpha: Hylicelloidea)
Author
Lambkin, Kevin J.
text
Zootaxa
2020
2020-06-12
4790
3
525
539
journal article
10.11646/zootaxa.4790.3.7
1175-5326
3891303
F011C13E-8E90-45D8-ABA3-B7D58582648A
Crosbella elongata
Evans, 1956
(
Figs 4–8
)
Crosbella elongata
Evans, 1956: 192
, fig. 5H;
Carpenter, 1992: 233
, fig. 150.1;
Martins-Neto
et al
., 2003: 253
;
Jell, 2004: 55
,
2 unnumbered figs.
Crosbella alata
Evans, 1956: 193
, fig. 5K;
Evans 1961: 15
;
Martins-Neto
et al
. 2003: 253
;
Jell, 2004: 55
,
syn. nov.
Mesocicadella venosa
Evans, 1956: 194
, fig. 5I;
Carpenter, 1992: 257
, fig. 163.2;
Martins-Neto
et al
., 2003: 253
;
Jell, 2004: 63
,
2 unnumbered figs.
syn. nov.
Mesocicadella punctata
Evans, 1961: 14
, fig. 1A;
Jell, 2004: 63
,
syn. nov.
(partim)
Triassoscelis anomala
:
Evans, 1961: 16
(specimen UQE1/1A).
Material
(all Mount Crosby).
Holotype
tegmen
UQC
2125,
holotype
tegmen and part hindwing of
C. alata
UQC
173/174,
paratype
hindwing of?
C. alata
UQC
1923/1924,
holotype
tegmen of
M. venosa
UQC
844,
holotype
tegmen of
M. punctata
QMF
3681a/b. 1841. Five additional tegmina:
UQE
1/1A, identified as
Triassoscelis anomala
by
Evans (1961)
;
ACC
: I.38a/b, I.39, I.49a/b, I.51.
Description.
Tegmen
c
.
19.7–20.5 mm
long (
holotype
20.5),
c
. 5.8–
c
.
6.4 mm
wide (
holotype
c
. 6.4), length/ width 3.2–3.4; apparently entirely darkly shaded (
Figs 7, 8
), although the counterparts of entirely darkly-shaded specimens have the vein margins pale, especially noticeable around the costal veinlets (
Fig. 4
).
Notes.
Evans distinguished his
C. alata
on its tegmen length of
14 mm
, ‘widely-spaced punctures’, and M and CuA
1
‘with accessory terminal veins’, compared to the type species with tegmen length of
12 mm
, ‘shallow pits’, M
1
simple, M
2
and M
3
with additional branches, and CuA ‘not preserved’. There is, however, some variation in tegmen length and considerable variation in the details of the apical venation of the species, and
C. alata
falls within this range. Its punctation is poorly preserved and scarcely detectable. Evans’ figure (1956, fig. 5K—compare with
Figs 5, 6
) does not show the three weak costal veinlets and several apical crossveins, and has CuA
1b
simple rather than branched. The
holotype
of
C. alata
also preserves what appears to be a small portion of the hindwing (
Fig. 6
). Its venation is similar although not exactly the same as the
paratype
specimen, UQC1923/1924, which Evans identified as the hindwing of
C. alata
. In the absence of venational congruence of this hindwing with the hindwing preserved in the
holotype
specimen, and confirmed association with the tegmen, UQC1923/1924 is therefore only tentatively referred to
C. elongata
.
Mesocicadella venosa
was separated on the presence of costal veinlets, accessory branches on RA and RP, and additional branching of CuA
1
, all of which match the venational scheme of
C. elongata
. Evans’ figure (1956, fig. 5I) does not show the multi-branched CuA
1
and lacks several apical crossveins. The
holotype
of
M. punctata
is again a typical specimen of
C. elongata
.
Evans (1961)
distinguished it on ‘the veins apically simple instead of being multi-branched’, but the apex of the specimen is not preserved and those post-nodal veins which are preserved match the branching pattern of
C. elongata
. The specimen preserves more of the multi-branched state of CuA than is shown in Evans’ figure (
Evans 1961
, fig. 1A). Specimen UQE1/1a, identified by
Evans (1961)
as
Triassoscelis anomala
, preserves only the central and antero-apical venation which matches that of the
holotype
of
C. elongata
.