Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae) Author Pall-Gergely, Barna https://orcid.org/0000-0002-6167-7221 Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary pallgergely2@gmail.com Author Hunyadi, Andras Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary Author Vermeulen, Jaap J. JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands Author Grego, Jozef Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia Author Sutcharit, Chirasak Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Reischuetz, Alexander Puechhaimgasse 52, A- 3580 Horn, Austria Author Dumrongrojwattana, Pongrat Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand Author Botta-Dukat, Zoltan Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary Author Oerstan, Aydin 12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA Author Fekete, Judit University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary Author Jochum, Adrienne https://orcid.org/0000-0002-6624-6412 Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany text ZooKeys 2023 2023-02-13 1147 1 177 http://dx.doi.org/10.3897/zookeys.1147.93824 journal article http://dx.doi.org/10.3897/zookeys.1147.93824 1313-2970-1147-1 9BB9881B0076473D8E53155D37CA1F50 FF2B6B317B505F9EA0E1000BDCD16CE7 Angustopila milium (Benson, 1853) Fig. 18 Cyclostoma Cyclostoma Milium Benson, 1853: 285. Hydrocena milium - W.T. Blanford 1864 : 464. (perhaps Georissa ); Godwin-Austen 1872 : 515, pl. 30, fig. 3. Cyathopoma milium - W.T. Blanford 1869 : 178. Georissa milium - Pfeiffer 1875-1876 : 292. Acmella milium - Gude 1921 : 362; Ramakrishna et al. 2010 : 111. Angustopila milium - Das et al. 2021 : 329; Preece et al. 2022 : 154, fig. 70A. Type locality. "in muscis arborum ad Musmai, prope Cherra-poonjee" (India, Meghalaya, Cherrapunji, approximate GPS coordinates: 25°16.6'N , 91°44'E ). Material examined. One syntype , possibly holotype , with no locality data (UMZC I.103805). An old label in unknown handwriting reads " Hydrocena milium Bs." and on the reverse side: "original shell described". Distribution. Known from a single shell only (Fig. 19 ). Figure 18. Angustopila milium (Benson, 1853) (syntype, UMZC I.103805). Photos: H. Taylor (NHM). Scale bar 1 mm. Figure 19. Distribution of Angustopila species. Remarks. Angustopila milium was described as a Cyclostoma (caenogastropod) species. Even Godwin-Austen (1872) was convinced that this was an operculate species when he examined Benson's type specimens. Gude (1914) considered it as juvenile Boysidia plicidens . However, the combination of the shell characters (small, conical shell, narrow, deep umbilicus, toothless, subcircular aperture, expanded aperture, sparsely arranged spiral striae) indicates that this species represents an adult hypselostomatid, which undoubtedly fits to the genus Angustopila , significantly extending the genus distribution. Here, we do not present a formal redescription because the type specimens could not be examined by SEM, and therefore, some important details cannot be seen. However, due to the large geographic distance between A. milium and other species discussed herein, it is highly unlikely that A. milium would be conspecific with any of them.