Neotypification of Drawida hattamimizu Hatai, 1930 (Annelida, Oligochaeta, Megadrili, Moniligastridae) as a model linking mtDNA (COI) sequences to an earthworm type, with a response to the ' Can of Worms' theory of cryptic species
Author
Blakemore, Robert
Hanyang Uni Seoul, Seoul, Korea, South
Author
Kupriyanova, Elena
,, Japan
Author
Grygier, Mark
Lake Biwa Museum, Kusatsu ,, Japan
text
ZooKeys
2010
2010-03-26
41
4
1
29
journal article
10.3897/zookeys.41.374
00f6e7e3-859e-422d-b9a5-51c458a7575a
1313–2970
576648
Type-species
:
Moniligaster deshayesi
Perrier, 1872
from
Sri Lanka
.
Family
diagnosis:
(from
Michaelsen 1900: 109
;
Stephenson 1923: 116
;
Gates 1962: 299
,
1972: 238
;
Blakemore 2008a
): Setae lumbricine. Prostomium prolobous. Male pores one or two pairs (on 10 or in 10/11 or 11/12 and 12/13) behind spermathecal pores (in 7/8 and/or 8/9) and in front of female pores (in 11/12 or near 12/13 or 13/14). Testis and male funnels intraseptal in sacs. Gizzards multiple (e.g.
2–12 in
region of segments 11–34) behind ovarian segment. Nephridia holoic. Clitellum unilayered (one cell thick). Ova macrolecithal (i.e., with large yolk-particles). Spermathecae with long tubular ducts. Last hearts in segment 9.
Distribution
:
Southeastern and eastern Asia, from
Sri Lanka
to eastern Siberia, also
Korea
,
Japan
, the
Philippines
, Borneo and Sumatra. Several species of
Drawida
have been reported outside the generic range, some of these later synonymized, but most moniligastrids are probably confined to their areas of origin in the Indo-Asian region (
Gates 1972
).