Review of Chilean Cyrtinae (Diptera: Acroceridae) with the Description of Three New Species and the First Record of Villalus inanis from Argentina Author Barahona-Segovia, Rodrigo M. Author Guzmán, Vicente Valdés Author Pañinao-Monsálvez, Laura text Zoological Studies 2021 2021-07-07 60 35 1 18 journal article 54167 10.6620/ZS.2021.60-35 dab15414-b59f-48a1-96ce-2515b2143001 PMC8652416 34963788 8070503 Holops virens Bigot 1878 : lxxii ( Figs. 13–14 , 20 ) Material examined : Syntype : “ Chile ” (A F UMO); other specimens: Chile : Atacama : Los Loros, 19.viii.1966 , Leg. C. Vivar (MNHNCL); Metropolitana : Cerro San Cristobal ( Stuardo 1980 ); Laguna Aculeo, 400 m , 22.viii.2014 , Leg. A. Ramírez (FRPC); Ñuble: Cerro Malalcura, San Fabián de Alico, VIII Reg. 1100 m . 11 X 2014 , Leg. A. Ramírez (FRPC); La Araucanía : Camping Pehuenco, Parque Nacional Nahuelbuta, 16.x2013 , Leg. F. Ramírez (FRPC). Type Locality : Chile ( Pape and Thompson 2013 ). Diagnosis : head and eyes black. Thorax and abdomen shiny green metallic color; dense and short whitish pilosity. Tibia and tarsi yellowish with short pilosity of same color. Wing hyalines. Description : Length: 5.1 mm ; width: 1.53 mm (head); 3.6 mm (thorax); 4.46 mm (abdomen); wing : 5.31 mm ; male : head : black; the size is 3/4 of the maximum height of the thorax; antenna inserted in the middle of the head; scape and pedicelum blackish with similar size; flagellum styliform three times longer than pedicelum and scape together; antennae approximately 1/4 shorter of maximum head length; eyes black with long and brown pilosity ( Figs. 10–11 ); three ocelli dark brownish with ocellar triangle black somewhat protuberant; occiput brownish with brown pilosity; mouth parts dark brownish; thorax : dark green scutum with long, dense and yellow-brownish pilosity, sparse but uniformly distributed ( Figs. 10–11 ); postocellar lobe dark green with long and yellowish pilosity; anterior spiracle yellowish; proepimeron, anepisternum, katepisternum, anepimeron, meron, katatergite, anatergite dark green with long and yellowish pilosity ( Fig. 11 ); scutellum dark green with whitish pilosity ( Fig. 11 ); legs : coxa, trochanters and femora are brownish excepting anterior part which is yellow; tibiae are yellowish; tarsi are brownish with dense brown pilosity; black claws ( Figs. 10–11 ); wings : smoky and with membranous appearance; all veins dark brown; R 4 and R 5 forked and outward separated for the half of length of each vein ( Fig. 20 ); M 1 , M 2 and M 3+4 not reaching the wing margin; M 1 shorter than M 2 ; cell r 4+5 elongated, with five sides and thin, with approximately more than the half of total length of R 2+3 vein; cell m 3 with 1/3 length of r 4+5 and connected directly with m-cu vein ( Fig. 20 ); halters brownish with apex somewhat yellowish; calypter rim blackish and interior part is brownish with dense and long brownish pilosity; Abdomen : elongated; slightly wider than thorax margins; fully rough; all tergites with dark green color and yellowish pilosity; all sclerite of greenish color ( Figs. 10–11 ); Genitalia : not detached. Figs. 9–12. Holops grezi sp. nov. : (9) dorsal view and (10) lateral view; Holops pullomen sp. nov. : (11) dorsal view and (12) lateral view. Scale bars = 1 mm. © 2021 Academia Sinica, Taiwan Distribution : from Atacama region to La Araucanía region ( Fig. 27 ). Remarks : Endemic species barely collected. The specific epithet “ virens ” means green (in English) or verde (in Spanish). This spider fly species is distributed from Coquimban province to Pehuén district in Maule province , both belong to central Chile subregion ( Morrone 2015 ). This species inhabits several vegetation types along its distribution (Andrés Ramírez com. pers.). Following Stuardo (1980) this species visits the flowers of Podanthus mitique Lindl. At same that H. frauenfeldi , H. virens could use dead branches of Cryptocarya alba (Molina) Looser and Podocarpus saligna D. Don to rest and watch potential competitors ( González et al. 2018 ). The species present high variability in color legs and length of R 4 , R 5 and M veins of wings (Shaun Winterton, pers. comm.). The Host is unknown.