The fifth family of the true crickets (Insecta: Orthoptera: Ensifera: Grylloidea), Oecanthidae defin. nov.: phylogenetic relationships and divergence times Author Campos, Lucas Denadai De Author Dias, Pedro Guilherme Barrios De Souza Author Audino, Jorge Alves Author Desutter-Grandcolas, Laure Author Nihei, Silvio Shigueo text Zoological Journal of the Linnean Society 2023 2022-11-02 197 4 1034 1077 http://dx.doi.org/10.1093/zoolinnean/zlac066 journal article 57261 10.1093/zoolinnean/zlac066 8c2036d0-8307-4b09-8c8b-c5eeea9d030a 0024-4082 7803427 4BB4333D-64F0-4485-9C2B-47546ECFE65F SUBFAMILY PODOSCIRTINAE SAUSSURE, 1878 DEFIN. NOV. Podoscirtites Saussure, 1878: 693 , Saussure, 1893: 257. Eneopterinae Kirby, 1906: 92 , Chopard, 1968: 365 , Chopard, 1969: 326. Vickery, 1977: 14, Otte & Alexander, 1983: 1–477 . Eneopteridae Kevan, 1977: 40 . Podoscirtidae Bruner 1916: 416 , Chopard, 1951: 491, Desutter, 1987: 224 , Desutter, 1988: 361 . Podoscirtinae Gorochov, 1986: 519 , Otte, Alexande & Cade, 1987: 448, Vasanth, 1993: 114, Otte, 1994: 72 , 86. Oshiro, 1995: 37, Yin, Haisheng & Liu, 1995: 213, Ingrisch, 1997: 48 , Gorochov, 2002: 303–350, Gorochov, 2003: 267-303, Gorochov, 2004: 33, Gorochov, 2005: 181– 208, Gorochov, 2006: 33–46, Gorochov, 2007: 237–289, Gorochov, 2008: 15, Gorochov, 2010: 205–245 , Gorochov, 2011: 216–270 , Gorochov, 2013: 15–58 , Gorochov, 2015: 31–41 , Anso, Jourdan & Desutter-Grandcolas, 2016: 30 , Desutter-Grandcolas, Anso & Jourdan, 2016: 436 , Gorochov, 2017: 11–106 , Gorochov, 2018: 77–121 , Rentz & Su, 2019: 124, Gorochov, 2020: 248 , Gorochov, 2021: 65 , Gorochov, 2021: 381–389 , Zheng et al ., 2021: 401. Type genus: Podoscirtus Serville, 1838 . Distribution: Worldwide. Diagnosis: Small to large-sized, body generally fusiform, intensely covered by bristles, with developed FWs, sometimes with stridulatory apparatus not developed. TIII with five or more outer and inner subapical spurs. Apex of dorsal valves of ovipositor serrulate laterally and ventrally, covering ventral valves laterally and sometimes ventrally ( Fig. 9B, C ). Male genitalia: apex of ectophallic fold bilobate; posterior projection of endophallus double (U-shaped) ( Fig. 9A ) when present; EndAp generally flattened laterally. Included supertribes and genera: Podoscirtidi Saussure, 1878 supertrib.nov. , Hapithidi Gorochov, 1986 supertrib. nov. Incertae sedis: Allopterites Cockerell, 1920 (fossil), Stenogryllodes Chopard, 1936 (fossil).