Taxonomic revision of the genus Amaranthus (Amaranthaceae) in Saudi Arabia
Author
Hassan, Walaa A.
Author
Al-Shaye, Najla A.
0000-0002-0447-8613
Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia & naaalshaye @ pnu. edu. sa; https: // orcid. org / 0000 - 0002 - 0447 - 8613
naaalshaye@pnu.edu.sa
Author
Alghamdi, Salma
0000-0001-7716-2233
Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia & saatalghamdi @ pnu. edu. sa; https: // orcid. org / 0000 - 0001 - 7716 - 2233
saatalghamdi@pnu.edu.sa
Author
Korany, Shereen M.
0000-0002-9355-3952
Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia & smkorany @ pnu. edu. as; https: // orcid. org / 0000 - 0002 - 9355 - 3952
smkorany@pnu.edu.as
Author
Iamonico, Duilio
0000-0001-5491-7568
Ce. R. S. I. Te. S., University of Rome Sapienza, viale XXIV Maggio 7, 04100 - Latina, Italy & duilio. iamonico @ uniroma 1. it; https: // orcid. org / 0000 - 0001 - 5491 - 7568 & Authors that contributed equally to the work
text
Phytotaxa
2022
2022-12-20
576
2
135
157
http://dx.doi.org/10.11646/phytotaxa.576.2.1
journal article
53324
10.11646/phytotaxa.576.2.1
1a9bd59f-7293-432f-b3c1-6df0bc659474
1179-3163
7461280
8.
Amaranthus spinosus
L
., Sp.
Pl. 2: 991. 1753
Type (
lectotype
designated by
Fawcett & Rendle 1914: 103
):—ASIA.
Habitat in Indiis
,
Herb. Linn. No.
1117.27 (LINN!, image of the lectotype available at http://linnean-online.org/11653/)
.
Description:—
Herbs 1–85(–120) dm tall, monoecious, annual (therophyte), rarely biennial. Stems usually erect, ± glabrous (sometimes sparsely pubescent in the inflorescence region), white to white-greenish, much branched. Leaves green (usually pale green), ovate, elliptic to spathulate (3.5–6.0 × 1.5–3.0 cm in the main axis, greater than the leaves on the branches), often with undulate margins (sometimes with a white marginal vein), apex obtuse and mucronate, base cuneate, glabrous (rarely pubescent on the veins), petioled (petiole 1.6–3.0 cm long). Synflorescences arranged in axillary glomerules, light green. Floral bracts greenish, ovate-lanceolate [(2.0–)3.0–4.0(–6.0) ×
0.3–0.6 mm
], 2(–2.5) times longer than the perianth, awned, margin entire, glabrous. Staminate flowers with 3 tepals, lanceolate; stamens 3. Pistillate flowers with 3 tepals, linear to lanceolate [0.9–1.1 × 0.3–0.4(–0.5) mm], with acute apex; stigmas 3. Fruit brownish-black, ellipsoidal [(1.2–)1.4–1.8 × 1.0–1.2(–1.4) mm], as long as or slightly longer than the perianth, rugose when dry, dehiscent. Seed lenticular [(0.8–)0.9–1.1(–1.3) mm in diameter], black to brownish-black.
Iconography:—
Beck (1909
: Tab. 297, figures 3–5);
Chaudhary (1998
: Plate.
127g
–j),
Bayón (2015: 296
, Figura 15).
Phenology:—
Flowering time June (
Al-Turki
et al.
2000
).
Habitat and elevation:—
Human-made habitat, about sea level (
Al-Turki
et al.
2000
).
Chromosome number:—
2n = 34 (
Al-Turki
et al.
2000: 341
), 68.
Alien status:—
Neophyte species native to Neotropics, it can be considered as casual in
Saudi Arabia
(see also
Chaudhary 1998
).
Occurrence in
Saudi Arabia
(
Fig. 10
):—
Bisha (
Abbas
et al.
2020
),
Jizan
(
Al-Turki
et al.
2000
, Aljieddani
et al.
2021). No finding was done during the filed surveys. Further researches are necessary to verify the distribution of
A. spinosus
in the country.