Revision of the spider genera Nilus O. Pickard-Cambridge 1876, Sphedanus Thorell 1877 and Dendrolycosa Doleschall 1859 (Araneae: Pisauridae) Author Jäger, Peter text Zootaxa 2011 3046 1 38 journal article 46245 10.5281/zenodo.278844 da2ca486-83fa-4e10-99eb-f3374d9a2a51 1175-5326 278844 Dendrolycosa bairdi spec. nov. Figs 84–95 , 167 : 9 (see also Figs 162–163, 165–166 ) Type material. Male holotype ( SMF ), L95, Laos , Champasak Province, Muang Bachieng, 38 street km E of Pakse, Ban Lak 38, That Fane, N 15°11´03.0´´ , E 106°07´36.9´´ , 952 m altitude, Bolaven plateau, coffee plantation, at day, by hand, P. Jäger & J. Martens leg. 15.III.2010 as subadult; adult in laboratory: IV.2010 . Female paratype ( SMF ), L95, Laos , Champasak Province, Muang Bachieng, 38 street km E of Pakse, Ban Lak 38, That Fane, N 15°11´03.0´´ , E 106°07´36.9´´ , 952 m altitude, Bolaven plateau, coffee plantation, at day, by hand, P. Jäger & J. Martens leg. 11.III.2010 . Additional material examined. 1 juvenile female ( SMF ), L95, Laos , Champasak Province, Muang Bachieng, 38 street km E of Pakse, Ban Lak 38, That Fane, N 15°11´03.0´´ , E 106°07´36.9´´ , 952 m altitude, Bolaven plateau, coffee plantation, vegetation and leaf litter, at night, by hand, P. Jäger & J. Martens leg. 19.III.2010 (together with type series of D. duckitti spec. nov. ). FIGURES 71–80. Dendrolycosa robusta (Thorell 1895) from Laos, Vientiane, Ban Dongmakkhai, Houay Nhang Conservation Forest, habitus, web and habitat (71 ventral view; 72–74, 77–78 dorsal view; 75 lateral view; 76 frontal view; 79 web in original habitat; 80 Original habitat). 71–73, 75–77 Adult female. 74, 78–79 Same specimen, but immature (two moults left to adult stage). Note differences between specimens preserved in ethanol (71–72) and live spiders. Etymology. The species is named in honour of Ian Baird (Madison, Wisconsin), the founder of the organisation “Global Association for People and Environment”, for his efforts to support local tribes in Laos and increase their standard of living; noun (name) in genitive case. Diagnosis. Medium sized spiders with body length 10.0– 15.1 mm with colour pattern A ( Figs 91–95 ). Male copulatory organ similar to that of Dendrolycosa robusta , but differing in following characters ( Figs 84–87 ): 1. Tibia shorter than cymbium, with broad RTA base in retrolateral view. 2. Median apophysis triangular with wide membranous base. 3. Conductor longer, i.e. arising more proximally and reaching further distally. Female copulatory organ similar to that of D. songi in having entire carina, but differing in following characters ( Figs 88–90 ): 1. Carina concave, fossae not as distinctly separated as in D. songi . 2. Middle field posteriorly broad, i.e. lateral lobes not covering field almost entirely as in D. songi . 3. Copulatory ducts broad, touching medially, extending laterally beyond spermathecal heads, covering spermathecal stalks and anterior bases of spermathecae. Description. Male ( holotype ). PL 4.3, PW 3.5, AW 2.0, OL 5.7, OW 3.4. Eye diametres: AME 0.22, ALE 0.21, PME 0.22, PLE 0.22. Eye interdistances: AME–AME 0.15, AME–ALE 0.04, PME–PME 0.19, PME–PLE 0.30, AME–PME 0.27, ALE–PLE 0.45, clypeus AME 0.23, clypeus ALE 0.25. Leg and pedipalpus measurements: pedipalpus 4.5 (1.5, 0.7, 0.7, -, 1.6); leg I 17.2 (4.7, 2.0, 1.6, 3.9, 3.7, 2.0); leg II 15.3 (4.4, 1.6, 3.9, 3.7, 1.7); leg III 13.8 (4.1, 1.4, 3.1, 3.5, 1.7); leg IV 17.3 (4.8, 1.7, 4.0, 4.8, 2.0); leg formula 4123. Spination: palp: femur 150, patella 110, tibia 1102; Femur I 336, II 338, III 535, IV 432; Patella I–IV 121, Tibia I 2128 , II 2228 , III–IV 2226 ; Metatarsus I 3027, II–IV 3037 (metatarsi I–III with, IV without distal median ventral spine). Chelicerae with 3 anterior (median largest) and 3 equally sized posterior teeth. Paired leg claws with c. 13 teeth. Copulatory organ as in diagnosis. RTA arising distally, tapering and distally bent in ventral view; with subdistal ventrad projection in retrolateral view. Cymbium with distinct prolateral and retrolateral bulge, with narrow tapering tip. Embolus filiform, arising from ventral tegulum, basal part visible in retrolateral view, distal part situated in conductor; tip as wide as distal part. Distal apophysis tapering, pointed, retrolaterad. Distal tegular projection bulgy, extending distinctly beyond prolateralcymbial margin. Subtegulum well visible in proximal part of alveolus ( Figs 84–87 ). Colour in ethanol ( Figs 91–92 ): yellowish to reddish brown with strongly contrasting pattern. Dorsal prosoma with 2 broad bright marginal bands, darker in posterior half. Eye region, fovea and triangular area in front dark. Sternum and ventral coxae without pattern. Gnathocoxae yellowish brown, labium reddish brown with white distal lip. Chelicerae reddish brown with basal humps, bright and longitudinal dark line in frontal view. Palp and legs with dark pattern, the latter most conspicuous in dorsal femora. Dorsal opisthosoma with characteristic dark patch in anterior half bordered by thin bright margins and a lighter brown folium, the latter darker and with undulating lateral margins in posterior half. Lateral opisthosoma brightly contrasting and with irregular patches. Ventral opisthosoma with 2 broad lateral and one narrow median indistinct longitudinal bands. Colour of live specimens more greyish brown, pattern on legs and body bright due to white hairs ( Figs 93, 95 ). FIGURES 81–83. Dendrolycosa songi (Zhang 2000) from China, Menglun (81 epigyne, ventral view; 82 vulva, dorsal view; 83 schematic course of internal duct system). Modified from Zhang (2000), not to scale. Female ( paratype ). PL 6.2, PW 4.9, AW 3.0, OL 8.9, OW 5.5. Eye diametres: AME 0.30, ALE 0.30, PME 0.30, PLE 0.32. Eye interdistances: AME–AME 0.18, AME–ALE 0.19, PME–PME 0.29, PME–PLE 0.46, AME– PME 0.32, ALE–PLE 0.61, clypeus AME 0.45, clypeus ALE 0.46. Leg and pedipalpus measurements: pedipalpus 6.6 (2.1, 1.1, 1.2, -, 2.2); leg I 19.1 (5.3, 2.5, 4.8, 4.5, 2.0); leg II 19.0 (5.5, 2.5, 4.7, 4.3, 2.0); leg III 15.4 (4.7, 1.8, 3.5, 3.7, 1.7); leg IV 19.2 (5.6, 2.0, 4.6, 4.9, 2.1); leg formula 4123. Spination: palp: femur 151, patella 110, tibia 1101, tarsus 1014; Femur I 335 (retrolateral ones short), II–III 535, IV 332; Patella I–IV 121, Tibia I–II 2128 (dorsal thin and short), III–IV 2226 ; Metatarsus I 3035, II 3027, III–IV 3037 (all metatarsi with distal ventral median spine). Chelicerae as in male. Palpal claw with 7–8 teeth, paired leg claws with c. 13 teeth. Copulatory organ as in diagnosis. Epigynal field wider than long, with two inconspicuous humps at anterior margin and two pairs of slit sense organs antero-laterally. Carina entire, with posterior pointed indentation. Fine wrinkles in anterior and posterior part of epigyne. Internal duct system with large copulatory ducts, the latter roughly half-moon shaped. Head of spermathecae distinct, their visible part as wide as long. Base of spermathecae round ( Figs 88–90 ). Colour in ethanol ( Fig. 94 ): as in male, but generally darker and pattern less contrasting. Dorsal prosoma dark reddish brown with 2 submarginal bright bands, with fovea dark. Labium and gnathocoxae dark reddish brown basally becoming pale yellow distally. Chelicerae dark reddish brown. Palp and legs dark reddish brown with obscure pattern especially on femora. Dorsal opisthosoma deep reddish brown with characteristic patch in anterior half, bordered by bright median line and broad reddish brown folium, the latter becoming darker in posterior half. Ventral opisthosoma dark yellowish brown with indistinct longitudinal lines. Colour of live specimens are probably shown in Figs 165–166 (specimens have not been caught and identified). Natural history. Webs were recorded from vegetation in natural and human modified habitats ( Figs 162–163 ). Distribution. Known only from the type locality.