Review oI the genus Ammatomus A. COSTA, 1859 (Hymenoptera, Crabronidae) Irom the Palearctic region with description oI Iour new species Author Schmid-Egger, Christian christian@bembix.de text Linzer biologische Beiträge 2019 2019-07-26 51 1 437 457 journal article 22915 10.5281/zenodo.3758403 8ca4d6b5-953c-443f-b6e5-fa12798918b0 0253-116X 3758403 Key to species of Ammatomus from the Palearctic region including Yemen Key to females 1 T1 large, not petiolate, abdomen without constriction between T1 and T2 ( figs 3 , 19). [TI at most as long as maximum width] ( A. aljahdhamii is somewhat intermediate, see description)..........................................................................................................................2 - T1 strongly petiolate, with distinct constriction between T1 and T2 ( fig. 4 ).......................3 2 Mesonotum, metanotum and sternites black. Tergal bands narrow, as large as length of metanotum ( fig. 1 ). 10 mm . Oman .......................... A. aljahdhamii SCHMID- EGGER nov.sp. - Mesonotum, metanotum and at least S2 with yellow bands or spot. Tergal bands at least 2x as large as length of metanotum ( fig. 19 ). [Mesosoma and abdomen with coarse punctation. Band of T1 and T2 medially largely interrupted. Propodeum without pubescence.]. 9-11 mm . Southeastern Europe to Central Asia .............................................. ................................................................................. Ammatomus rogenhoferi HANDLIRSCH 3 Hindtarsomeres I-III apically black ( fig. 22 ). S2 with large yellow band (cf. fig. 24 ), except in A. coarctatus ........................................................................................................4 - Hindtarsomeres I-III all yellow ( fig. 6 ). S2 with lateral triangular spots.............................7 4 T1 with lateral yellow spots. T1 shorter than maximum width ( fig. 8 ). S2 with lateral triangular spots. [Punctation of mesoscutum and other parts of mesosoma dense, most interspaces smaller than punctures. T1 black apart from yellow spots.]. Southern Europe to Central Asia ..................................................... Ammatomus coarctatus SPINOLA - T1 with continuous yellow band ( fig. 7 ). T1 at least 1.3x ...................................................5 5 T1 1,3x - 1.4x. [T1 red, with yellow apical band]. Algeria ................................................... ............................................................................ Ammatomus saharae (HANDLƖRSCH̹ 1895) - T1 at least 1,5x ....................................................................................................................6 6 Yellow band on T1 medially not emarginated. Transparent apical margin of T1, T3 and T4 maximal half as wide as yellow band ( fig. 21 ). Scutellum with large yellow band, mesoscutum laterally with large yellow band in whole length ( fig. 20 ). [Pubescence of propodeal declivity longer and sparse, compared to A. grandcourti . T1 apart from yellow band always red,]. Turkey to Central Asia; Oman ..................................................... ................................................................................ Ammatomus rufonodis RADOSZKOWSKI - Yellow band on T1 medially emarginated. Transparent apical margin of T1, T3 and T4 as wide as yellow band medially ( fig. 10 ). Scutellum black, yellow band of mesoscutum laterally in most specimens restricted to apical third ( fig. 9 ). [T1 apart from yellow band, red or black]. Israel . Arabia, Sudan ......................................................... ................................................................ Ammatomus grandcourti SCHMID- EGGER nov.sp. 7 Hind claw and hindtarsomere V yellow, at most with some brownish parts ( fig. 6 ). Lateral carina of T1 reaches apical depression of T1 ( fig. 5 ). Central Asia to North Africa ....................................................................... Ammatomus asiaticus RADOSZKOWSKI - Hind claws dark brown or black. At least apical half of hindtarsomere V evenly black, only apically brown in A. wathabensis . Lateral carina of T1 ending in basal half of tergite or shorter ..................................................................................................................8 8 T1 apically without clear transparent lamella, the apical margin has the same colour as disc or is darker, or somewhat paler red than disc. T1 apically with two narrow lateral spots or a narrow band; spots pale yellow, in a distance of about 1.5x diameter of hindocellar diameter apart from hind margin ( fig. 36 ). T 2 in most specimens red (apart from apical yellow band). Claws and apical half of hindtarsomere V brownish. Punctation of mesoscutum finer and interspaces larger, compared with punctation of mesopleuron ( fig. 38 ). [Lower mesopleuron with fine and dense silver pubescence. T1 1,2x] ........................................................ Ammatomus wathabensis SCHMID- EGGER nov.sp. - T1 apically with distinct transparent margin, and with yellow band or large lateral spots between margin and disc of tergite; remaining part of tergite red or black ( fig 32 ). T2 black. apart from apical yellow band. Claws and apical half of hindtarsomere V black. Punctation of mesoscutum similar to that of mesopleuron ( fig. 31 ) ....................................9 9 T1 1.3x - 1.4x ( fig. 32 ) ......................................................................................................10 - T1 shorter, at most 1.2x ( fig. 16 ). [Disc of T1 black]........................................................11 10 Mesoscutum with dense punctation, most punctures less than a diameter apart ( fig. 31 ) (however, puncture is irregular, and there are also larger interspaces). Body colour whitish yellow ( fig. 29 ). Disc of T1 red. Arabia, Iran , Israel ?............................................... .................................................................. Ammatomus vanharteni SCHMID- EGGER nov.sp. - Mesoscutum with sparse punctation, most punctures more than a diameter apart (fig. 50). Body colour lemon yellow (fig. 47). Disc of T1 black. Russia , Central Asia ................ .................................................................. Ammatomus nikolajevskii (GUSSAKOVSKƖJ̹ 1928) 11 Body colour lemon yellow. T1 with continuous yellow band ( fig. 14 ). T1 1x ( fig. 16 ). North Africa .............................................................. Ammatomus mesostenus HANDLIRSCH - Body colour whitish yellow. T1 with large lateral spots, space between spots as large as basal diameter of hindtibia ( fig. 41 ). T1 1.2x ( fig. 42 ). Yemen ............................................. ................................................................. Ammatomus yemenensis SCHMID- EGGER nov.sp. Key to males The male of A. aljahdhamii is undescribed. 1 T1 large, not petiolate, abdomen without constriction between T1 and T2 (cf. fig. 19 ). [T1 at most 1x. Mesosoma and abdomen with coarse punctation. Yellow bands on T1 and T2 medially interrupted. Propodeum without pubescence]. 8-10 mm . Southeastern Europe to Central Asia (if from Oman , cf. A. aljahdhamii ) .................................................. ................................................................................. Ammatomus rogenhoferi HANDLIRSCH - T1 strongly petiolate, with distinct constriction between T1 and T2. ( fig. 12 )....................2 2 Hindtarsomeres I-III apically black ( fig. 22 ). S2 with large yellow band, except A. coarctatus ( fig. 24 )..............................................................................................................3 - Hindtarsomeres I-III all yellow. S2with lateral triangular spots..........................................7 3 T1 with lateral yellow spots. T1 short, 1.3x (cf. fig. 8 ). S2 with lateral triangular yellow spots. [Punctation of mesoscutum and other parts of mesosoma dense, most interspaces smaller than punctures. T1 black apart from yellow spots]. Southern Europe to Central Asia ................................................................................... Ammatomus coarctatus SPINOLA - T1 and S2 with continuous yellow band. T1 at least 1.4x ...................................................4 4 T1 1.4x ( fig. 4 ). [T1 red with apical yellow band, medially emarginated, apical transparent margin of T1 as large as yellow band medially]. Algeria ................................... ............................................................................ Ammatomus saharae (HANDLƖRSCH̹ 1895) - T1 at least 1.6x ( fig. 23 ) [if shorter, not from Algeria ]........................................................6 5 T1 1.6x. lemmon yellow. Band on T1 medially not emarginated. Transparent apical margin of T1, T3 and T4 maximal half as wide as yellow band (in males from Oman is T1 longer, and yellow band of T1 narrower, see description) ( fig. 23, 26 ). Scutellum with large yellow band, mesoscutum laterally with large yellow band in whole length. [Pubescence of propodeal declivity longer and sparse, compared to A. grandcourti . T1 apart from yellow band always red,]. Turkey to Central Asia, Oman ................................... ................................................................................ Ammatomus rufonodis RADOSZKOWSKI - T1 1.7-1.8x. Tergal bands whitish-yellow. Band on T1 medially emarginated ( fig. 12 ). Transparent apical margin of T1, T3 and T4 as wide as yellow band medially ( fig. 13 ). Scutellum black, yellow band of mesoscutum laterally in most specimens restricted to apical third. [Pubescene of propodeal declivity shorter and very dense, compared to A. rufonodis . T1 apart from yellow band. red or black]. Israel , Arabia, Sudan .......................... ................................................................ Ammatomus grandcourti SCHMID- EGGER nov.sp. 6 Hind claw and hindtarsomere V yellow, at most with some brownish parts ( fig. 6 ). Lateral carina of T1 reaches apical depression of T1 ( fig. 5 ). Central Asia to North Africa ....................................................................... Ammatomus asiaticus RADOSZKOWSKI - Hind claws dark brown or black. At least apical half of hindtarsomere V evenly black, only apically brown in A. wathabensis . Lateral carina of T1 ending in basal half of T1 or is shorter..........................................................................................................................7 7 T1 apically without clear transparent lamella, the apical margin has the same colour as disc or is darker, or somewhat paler red than disc, apically with two narrow lateral spots or a narrow band; spots pale yellow, in a distance of about 1.5x diameter of hindocellar diameter from hind margin ( fig. 40 ). T 2 in most specimens red ( fig 39 ), (apart from apical yellow band). Claws and apical half of hindtarsomere V brownish.. [Lower mesopleuron with fine and dense silver pubescence. T1 1.4x] ................................. ................................................................ Ammatomus wathabensis SCHMID- EGGER nov.sp. - T1 apically with distinct transparent margin, and with yellow band or large lateral spots between margin and disc of tergite; remaining part of tergite red or black ( fig. 34 , 52). T2 black. apart from apical yellow band. Claws and apical half of hindtarsomere V black....................................................................................................................................8 8 T1 1,5x - 1.6.x (fig. 52, 53). [Body colour lemon yellow. Punctation of mesonotum in direct comparisation finer and more sparse as in remaining species]. Russia , Central Asia ........................................................... Ammatomus nikolajevskii (GUSSAKOVSKƖJ̹ 1928) - T1 at least 1.7x ( fig. 18 ) ......................................................................................................9 9 T1 red apart from yellow apical band ( fig. 34 ). [Scutellum in most specimens partly or all yellow. Body colour whithis yellow]. Arabia, Iran , Israel ? .............................................. ........................................................... Ammatomus vanharteni SCHMID- EGGER new species - T1 black apart from yellow apical band ............................................................................10 10 Body colour lemon yellow. T1 1.8x, slender ( fig. 18 ). Scutellum may be touched by yellow. North Africa. ................................................ Ammatomus mesostenus HANDLIRSCH - Body colour pale yellow. T1 1.7x, apically somewhat bulged ( fig. 46 ). Scutellum black. Yemen ........................................... Ammatomus yemenensis SCHMID- EGGER nov.sp.