Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species Author Toda, Masanori J. Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan Author Bänziger, Hans Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034 Author Sati, Pradeep C. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575 Author Fartyal, Rajendra S. Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263 Author Suwito, Awit Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com Author Katoh, Toru Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545 text Zootaxa 2020 2020-06-08 4789 1 1 54 journal article 21673 10.11646/zootaxa.4789.1.1 c12aaed8-264c-459c-b1aa-09eaeb686670 1175-5326 3884610 311A27A2-A053-4D20-A43E-EF57DBE55166 28) Phortica ( Ashima ) nigrifoliiseta (Takada, Momma & Shima, 1973) Amiota ( Phortica ) nigrifoliiseta Takada, Momma & Shima, 1973: 74 . Phortica ( Phortica ) nigrifoliiseta : Chen et al ., 2007: 25 . Phortica ( Ashima ) nigrifoliiseta : Chen & Máca, 2012: 507 . Diagnosis. Surstylus with approximately 8 apically blunt, peg-like teeth (ch.37-0); both anterior postgonites basally with strongly sclerotized, large flaps asymmetric in shape. Supplementary description (not repeating characters common to P . spinosa ). Supracervical setae 27–29. Arista with arb = 2/0. Cibarial, medial sensilla approximately 9 per side; posterior sensilla approximately 6 per side. Abdominal sternite III wider than long (ch.22-1). Pregonite with sensilla on submedial portion (ch.52-1). Posterior postgonite only on right side of aedeagal sheath; left, anterior postgonite apically strongly sclerotized and bifurcated, but right one not. Specimen examined. Malaysia : holotype ♂ , Silau Silau Trail , Mt. Kinabalu , Sabah , 27.vi.1972 , E. Momma leg. ( SEHU ) . Distribution. Borneo ( Sabah ).