Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America
Author
Smith, Andrew B. T.
Author
Evans, Arthur V.
text
Zootaxa
2018
2018-09-05
4471
2
279
308
journal article
29479
10.11646/zootaxa.4471.2.3
694f85ad-7913-43ea-89b9-8eb4e20f0821
1175-5326
1439674
CE4C6E73-09CB-4E74-91C5-771B3696DC52
Ulata argentina
Saylor, 1945
Figs. 61–65
,
74
.
Original combination
.
Ulata argentina
Saylor, 1945
: 442
.
Type
locality
.
Pucará
,
Lago Lacar
,
Neuquén
,
Argentina
(approximately
40.16°S
,
71.63°W
)
.
The original
type
locality was “
Bariloche
,
Rio Negro
,
Argentina
” (
Saylor 1945
).
FIGURES 54–56.
Dihymenonyx herrerai
Gutiérrez, 1949
. 54, dorsal habitus; 55, oblique habitus; 56, head and pronotum.
Type series
. Neotype at CMNC labeled a) “ARGENTINA / NEUQUEN / Lago Lacar / Pucará / Shajovskogleg. / Coll. Martínez.- / NOV.-964” (handwritten), b) “H. & A. HOWDEN / COLLECTION /
ex
. A.Martinez coll.” (typeset with black border), c) “
Ulata
♂ /
argentina
/ Saylor / A. MARTINEZ-DET. 1965” (handwritten and
typeset), d) “Southern Neotropical Scarabs / database #
AS
2615414 /
Ulata argentina
/
Saylor, 1945
♂
/ DET: A.B.T.
SMITH
2 008” (typeset with black border), e) “
ULATA
/
ARGENTINA
SAYLOR, 1945
♂
/
NEOTYPE
” (handwritten and typeset with black border on red paper).
Neotype
here designated
.
Saylor (1945)
stated that this species was described using only the
holotype
, which was presumably deposited in his personal collection (although no collection deposition was explicitly stated it was Saylor’s common practice to keep
holotypes
in his personal collection). Saylor’s personal collection was donated to the
CASC
in 1963 (
Ratcliffe 2016
), but this institution has no record of the
Ulata argentina
holotype
(the rest of Saylor’s primary
types
are catalogued and kept in the
type
collection). The first author carefully searched the entire
CASC
scarab collection for the
holotype
, but found nothing. Therefore, we conclude that the
holotype
has been lost. Due to the historic instability of the classification of this taxon, the
neotype
is designated with the express purpose of clarifying the taxonomic status of this species and genus. The
neotype
is consistent with the original description and originated from relatively close to the original
type
locality (less than
80 km
distance).
FIGURES 61–65.
Ulata argentina
Saylor, 1945
neotype. 61, dorsal habitus; 62, oblique habitus; 63, lateral male genitalia; 64, head and pronotum; 65, parameres.
Distribution
.
Fig. 74
.
Argentina
:
Neuquén
,
Río Negro
;
Chile
: V
Región de Valparaíso
,
Región Metropolitana de Santiago
, VIII
Región del Biobío
, IX
Región de la Araucanía
(based on the original
type
locality and specimens from AMNH, BMNH, CMNC, FMNH, TMSA, UMCE, VMDM).