Phylogenetic systematics of the enigmatic genus Horologion Valentine, 1932 (Coleoptera, Carabidae, Trechinae, Horologionini), with description of a new species from Bath County, Virginia Author Harden, Curt W. https://orcid.org/0000-0002-5226-8725 Department of Plant and Environmental Sciences, 277 Poole Agricultural Center, Clemson University, Clemson, South Carolina 29634, USA c_har@fastmail.com Author Davidson, Robert L. Department of Invertebrate Zoology, Carnegie Museum of Natural History, 4400 Forbes Avenue, Pittsburgh, Pennsylvania 15213, USA Author Malabad, Thomas E. Virginia Department of Conservation and Recreation, Division of Natural Heritage, Karst Program, 16 th Floor, 600 East Main Street, Richmond, Virginia, USA Author Caterino, Michael S. https://orcid.org/0000-0002-2597-5707 Department of Plant and Environmental Sciences, 277 Poole Agricultural Center, Clemson University, Clemson, South Carolina 29634, USA Author Maddison, David R. https://orcid.org/0000-0002-7152-3824 Department of Integrative Biology, Oregon State University, Corvallis, Oregon 97331, USA text Subterranean Biology 2024 2024-01-08 48 1 49 http://dx.doi.org/10.3897/subtbiol.48.114404 journal article http://dx.doi.org/10.3897/subtbiol.48.114404 1314-2615-48-1 989B4A14F7D54805822A2181DE5223A4 768E4D6A499F5E8DB5FA59ACA071AC36 Horologionini Jeannel, 1949 Horologionidae Jeannel, 1949: 91. Type genus: Horologion Valentine, 1932. Horologionini : Barr (1964 : 1). Horologionina : Bousquet (2012 : 711); Ortuno and Arillo (2015 : 584), Maddison et al. (2019 : 171). Diagnosis. The tribe consists of a single genus, Horologion , described in detail below. Members of the tribe possess characteristics of the supertribe Trechitae and several character states that, in combination, distinguish them from other tribes of Carabidae : eyes absent, frontal furrows weakly impressed, penultimate maxillary palpomeres glabrous, apical maxillary palpomeres elongate and narrow, pronotal lateral margins and hind angles absent, elytral humeri with carinate shelf, elytron with a single discal setigerous puncture, apical recurrent stria absent, mesoventrite bell-shaped and extended anteriorly, mesocoxae and metacoxae separated by processes of mesoventrite and metaventrite, all surfaces of external integument except for mouthparts and abdominal tergites generally setose, median lobe of aedeagus open dorsally, spermatheca with attached gland, female genital segment subtriangular, gonocoxites slender and largely glabrous.