Do circum-Antarctic species exist in peracarid Amphipoda? A case study in the genus Epimeria Costa, 1851 (Crustacea, Peracarida, Epimeriidae)
Author
Loerz, Anne
,, New Zealand
Author
Maas, Elizabeth
National Institute of Water and Atmospheric Research, Wellington ,, New Zealand
Author
Linse, Katrin
British Antarctic Survey, Cambridge, United Kingdom
Author
Coleman, Charles Oliver
Museum für Naturkunde Berlin, Berlin, Germany
text
ZooKeys
2009
2009-08-26
18
91
128
journal article
10.3897/zookeys.18.103
4afb9953-2445-42e2-97c4-566dee230136
1313–2970
576497
3A6234A8-F3A5-4F43-B4FB-89722D121684
Genus
Epimeria
Costa
,
1851
in Hope, 1851
?
Vertumnus
White, 1847: 89
[
nomen nudum
].
Epimeria
Costa
, 1851: 24
[
nomen nudum
].–
Costa
in Hope 1851: 46. –
Karaman and Barnard 1979: 108
.–
Watling and Holman 1980: 642
.–
Coleman 2007: 31
.
Pseudepimeria
Chevreux, 1912: 9
(
type
species
Pseudepimeria grandirostris
Chevreux, 1912
; original designation).
Subepimeria
Bellan-Santini, 1972b: 225
(
type
species
Subepimeria geodesiae
Bellan- Santini, 1972; original designation).
Epimeriella
.–
Walker 1906: 17
.–
Karaman and Barnard 1979: 107
.–
Coleman 2007: 56
(
type
species
Epimeriella macronyx
Walker, 1906
; by monotypy). syn. n.
Type
species.
Epimeria tricristata
Costa
,
1851
in
Hope
, 1851; by monotypy
.
Remarks.
The most recent family diagnoses for the
Epimeriidae
is that of
Coleman (2007)
,
Coleman and Barnard (1991)
, and
Barnard and Karaman (1991)
.
Figure 1.
Maximum parsimony topology (length 1599, CI 0.33, RI 0,6815). Branch support values are given: jackknife above, parsimony / likelihood bootstrap below branches. Sequences obtained in this study are marked in bold. Th e scale bar gives the number of nucleotide substitutions per branch length.
Based on the genetic data presented in the following and a detailed morphological evaluation of the weak morphological separation criteria between
Epimeriella
and
Epimeria
we herewith synonymize
Epimeriella
with
Epimeria
. Since we transfer the species
Epimeriella macronyx
Walker, 1906
;
Epimeriella scabrosa
Barnard, 1930
;
Epimeriella truncata
Andres, 1985
;
Epimeriella victoria
(
Hurley, 1957
a)
and
Epimeriella walkeri
Barnard, 1930
to the genus
Epimeria
, we herewith give a new genus diagnosis.
The diagnosis has been broadened from that given by
Barnard and Karaman (1991)
since it now included characters formerly predominantly occurring in the genus
Epimeriella
such as a smooth body and the laminar, none triturative pars molaris.
Diagnosis.
Body covered with teeth or processes or body poorly armed, almost smooth. Antenna 1 peduncular article 2 shorter than 1. Accessory flagellum present or absent. Mouthparts projecting quadrately. Upper lip incised or almost entire; epistome not very broad. Mandibular incisor ordinary, toothed, setal row present; molar blunt, strong, triturative or simple, conical or laminar. Lower lip inner lobes absent, outer lobes relatively broad. Hypopharyngeal gap sometimes widened. Maxilla 1 palp 2-articulate, article 2 ordinary. Maxilla 2 inner plate without facial row of setae. Maxilliped inner plate narrower but as long as outer plate, latter elongate; palp article 2 narrow and unproduced; palp article 4 well developed, unguiform or serrate. Coxae 1–4 progressively longer; coxae 4–5 forming ventral arc; coxa 4 long, polycuspidate. Gnathopods alike, articles 5–6 elongate, subchelate (typical), sometimes simple. Telson incised or cleft.
After synonymising
Epimeriella
with
Epimeria
,
the genus
Epimera
now includes 46 species:
Epimeria annabellae
Coleman, 1994
;
Epimeria bathyalis
Wakabara and Serejo, 1999
;
Epimeria bispinosa
Ledoyer, 1986
;
Epimeria bruuni
Barnard, 1961
;
Epimeria cora
Barnard, 1971
;
Epimeria concordia
Griffiths, 1977
;
Epimeria cornigera
(J.C.
Fabricius, 1779
)
;
Epimeria extensa
Andres, 1985
;
Epimeria georgiana
Schellenberg, 1931
;
Epimeria glaucosa
Barnard, 1961
;
Epimeria grandirostris
(
Chevreux, 1912
)
;
Epimeria heldi
Coleman CO (1998a)
;
Epimeria horsti
Lörz, 2008
;
Epimeria inermis
Walker, 1903
;
Epimeria intermedia
Schellenberg, 1931
;
Epimeria longispinosa
Barnard, 1916
;
Epimeria loricata
G.O.
Sars, 1879
;
Epimeria macrodonta
Walker, 1906
;
Epimeria macronyx
(
Walker, 1906
)
,
comb. n.
;
Epimeria monodon
Stephensen, 1947
;
Epimeria obtusa
Watling, 1981
;
Epimeria oxicarinata
Coleman, 1990
;
Epimeria pacifica
Gurjanova, 1955
;
Epimeria parasitica
(M. Sars, 1858)
;
Epimeria pelagica
Birstein and M. Vinogradov, 1958
;
Epimeria pulchra
Coleman, 1990
;
Epimeria puncticulata
Barnard, 1930
;
Epimeria reoproi
Lörz and Coleman, 2001
;
Epimeria rimicarinata
Watling and Holman, 1980
;
Epimeria robusta
Barnard, 1930
;
Epimeria robustoides
Lörz & Coleman, 2009
,
sp. n.
;
Epimeria rotunda
Wakabara & Serejo, 1999
;
Epimeria rubrieques
De Broyer & Klages, 1991
;
Epimeria scabrosa
(
Barnard, 1930
)
,
comb. n.
;
Epimeria schiaparelli
Lörz, Maas, Linse and Fenwick 2007
;
Epimeria semiarmata
Barnard, 1916
;
Epimeria similis
Chevreux, 1912
;
Epimeria subcarinata
Nagata, 1963
;
Epimeria tuberculata
G.O.
Sars, 1895
;
Epimeria truncata
(
Andres, 1985
)
,
comb. n.
;
Epimeria ultraspinosa
Wakabara and Serejo, 1999
;
Epimeria vaderi
Coleman CO (1998b)
;
Epimeria victoria
(
Hurley, 1957
)
,
comb. n.
;
Epimeria walkeri
(
Barnard, 1930
)
,
comb. n.
;
Epimeria yaquinae
McCain, 1971
.