An update of morphological and distributional data of the genus Patrera Simon (Araneae: Anyphaenidae: Anyphaeninae) with the description of twenty-five new species from Colombia
Author
Martínez, Leonel
0000-0002-4166-0561
leonelmarbio@gmail.com
Author
Brescovit, Antonio D.
0000-0002-1511-5324
antonio.brescovit@butantan.gov.br
Author
Villarreal, Eduardo
0000-0003-1303-2573
leonelmarbio@gmail.com
Author
Oliveira, Luiz Fernando M.
0000-0002-4054-4681
text
Zootaxa
2021
2021-01-19
4914
1
1
64
journal article
8672
10.5281/zenodo.4468383
5c5c8aad-73f1-4a26-bdca-d1d02ba3fec1
4468383
Patrera ramirezi
new species
Figures 12A–G
;
30
Type material.
Male
holotype
from
Salto del Tequendama Falls
(
4°35′10″N
;
74°18′0″W
),
Soacha
,
Cundinamarca
,
Colombia
,
25.VII.1967
,
P. & B. Wygodzinsky
leg. (
AMNH
).
Paratype
: female same data as holotype (
AMNH
).
Male
from
Reserva Natural El Secreto
,
Saragoa
(
10°53′42.5″N
;
73°59′58.6″W
),
2120m
,
Boyacá
,
Colombia
,
13.X.2001
,
C. Zudiria
leg. (ICN-Ar-9667)
.
Other material examined.
COLOMBIA
,
Boyacá
:
Iguape
,
Quebrada Carrizal
, (
5°25′12″N
;
73°27′24″W
),
3360m
, ♁1,
17.VIII-15.IX.2000
,
P. Reina
leg. (ICN-Ar-10612)
.
Cundinamarca
:
Choachí
,
30.IV.2009
, ♁1
D. Luna
leg. (ICN-Ar-6037)
.
Etymology.
The specific epithet is a patronym in honor of Martín Ramírez, who has greatly contributed to taxonomic and systematic studies of Neotropical spiders.
FIGURE 12A–D.
Patrera ramirezi
n. sp.
Male: (AMNH): A habitus, dorsal view; C right palp, retroventral view; D palp, ventral view; E palp, retrolateral view. Female (AMNH): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view. Abbreviations: dps, dorsal projection of the subtegulum; FD, fertilization ducts; SR, seminal receptacles. Scale bars: A–B: 2.16mm; C:1mm; D–E: 0.66mm; F–G: 0.25mm.
Diagnosis.
Males of
Patrera ramirezi
n. sp.
can be differentiated from those of the remaining species of the genus by their long retrolateral tibial apophysis, with small dorsal projection (
Fig. 12
C–E). Females are distinguished by their epigynum with exceptionally wide, transversal hood, and transversal copulatory ducts with small seminal receptacles, anteriorly positioned (
Fig. 12
F–G).
Description.
Male
(
Holotype
, AMNH). Cephalothorax and legs brown-reddish (
Fig. 12A
). Abdomen dorsally gray with dark spots from median to posterior area; ventrally white (
Fig. 12A
). Total length 5.1, carapace length 2.6, width 2.02. Clypeus height 0.08. Eye diameters and interdistances: AME 0.06, ALE 0.10, PME 0.10, PLE 0.06; AME–AME 0.06, AME–ALE 0.06, PME–PME 0.12, PME–PLE 0.08, ALE–PLE 0.02. Chelicerae 0.72 long, four promarginal teeth, six retromarginal denticles. Leg measurements: leg I—femur 2.9/ patella 1.0/ tibia 3.0/ metatarsus 2.7/ tarsus 1.3/ total 10.9; II—2.6/ 1.0/ 1.62/ 2.8/ 1.05/ 9.07; III—1.8/ 0.8/ 1.4/ 1.6/ 0.7/ 6.3; IV—2.63/ 0.8/ 2.15/ 2.8/ 0.9/ 9.28. Leg spination: I—tibia v2-2-0, metatarsus v2-2-0, p1-1-0, r1-1-0; II—tibia v2-2-0, metatarsus v2-2-0, p1-1-0, r1-1-0; III—tibia v2-2-1, p0-1-1, r0-1-1; IV—tibia v2-2-1, p0-1-1, r0-1-1; Abdomen: length 2.5, epigastric furrow 0.9 from tracheal spiracle, spiracle 1.0 from base of spinnerets. Palp: retrolateral tibial apophysis long, ventrally projected; cymbium longer than tibia; subtegulum longer than wide with flattened and apically subrounded distal projection; tegulum longer than wide; median apophysis laminar, short and medially situated; embolus long, apically filiform with narrow base and basally inserted on the tegulum (
Fig. 12
C–E).
Female
(AMNH,
Paratype
). Coloration as in the male (
Fig. 12B
). Total length 5.8, carapace length 2.9, width 2.1. Clypeus height 0.10. Eye diameters and interdistances: AME 0.06, ALE 0.12, PME 0.14, PLE 0.12; AME– AME 0.04, AME–ALE 0.06, PME–PME 0.16, PME–PLE 0.10, ALE–PLE 0.04. Chelicerae 1.0 long, four promarginal teeth, five retromarginal denticles. Leg measurements: leg I—femur 2.52/ patella 0.9/ tibia 2.5/ metatarsus 2.1/ tarsus 1.0/ total 9.02; II—2.45/ 0.9/ 2.3/ 2.1/ 0.9/ 8.65; III—1.8/ 0.9/ 1.4/ 1.6/ 0.7/ 6.4; IV—2.5/ 0.9/ 2.0/ 2.6/ 0.9/ 8.9. Leg spination: I—tibia v2-2-0, metatarsus p1-1-0, r1-1-0; II—tibia v2-2-0, metatarsus v2-2-0, p1-1-0, r1-1-0; III—tibia p0-1-1, r0-1-1; IV—tibia v2-2-1, p0-1-1, r0-1-1; Abdomen: length 2.85, epigastric furrow 0.8 from tracheal spiracle, spiracle 1.0 from base of spinnerets. Epigynum: hood transversal, very wide and posteriorly projected; lateral borders weakly sclerotized, parallel, wide and apically rounded; atrium long and ample; internally with copulatory ducts long and wide; seminal receptacles small, rounded and anteriorly situated on the copulatory ducts; spermathecae large, oval and anteriorly positioned; fertilization ducts almost as long as spermathecae (
Fig. 12
F–G).
Variation.
Males (n=3): total length: 6.83–7.33; carapace length: 3.07–3.44; femur I length 2.9–3.68.
Distribution.
Known from
Boyacá
and
Cundinamarca
departments
Colombia
(
Fig. 30
).