The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea)
Author
Shear, William A.
Author
Richart, Casey H.
Author
Wong, Victoria L.
text
Zootaxa
2020
2020-03-19
4753
1
1
78
journal article
21316
10.11646/zootaxa.4753.1.1
f7974bcf-11e7-4d1b-b09d-530dc2dbd54f
1175-5326
3983782
AA9F66B3-EF8C-4F6B-8F35-0BCBEE5122ED
Complicatella complicata
(
Shear, 1974
)
,
new combination
Bollmanella complicata
Shear 1974
, p. 146
.
Figs. 192–193
FIGS. 190–193.
Brunsonia chelanomagna
and
Complicatella complicata
.
Figs. 190, 191.
Brunsonia chelanomagna
.
Fig. 190. Anterior gonopods, posterior view. Fig. 191. Left posterior gonopod coxite, posterior view. Figs. 192–193.
Complicatella complicata
. Fig. 192. Left anterior gonopod, posterior view. Fig. 193. Posterior gonopod coxites, posterior view.
New Records
:
WASHINGTON:
Thurston Co.
,
Black Lake
and
Belmore Road
at 66
th
Avenue
,
Tumwater
, elev.
60 m
.
,
46.9889°N
, -
122.9659°W
,
19 November 2003
,
W. Leonard
, mm;
Hospital Creek
, above confluence with
Skookumchuck River
, 5 horiz mi S, 3 horiz mi
E of Vail
, elev.
245 m
.
,
46.7732°N
, -
122.5855°W
,
15 December 2003
,
W. Leonard
,
K. McAllister
, mm ff;
Rainier Road SE
, 3.0 mi
NW of Military Road
SE,
Fort Lewis Military Reservation
, elev. ca.
100 m
.
, ca.
46.9351°N
, -
122.7600°W
,
29 November 2003
,
W. Leonard
, m;
The Evergreen State College
campus,
Beach Trail
N of
Driftwood Road
,
Olympia
, elev.
30 m
.
,
47.0791°N
, -
122.9776°W
,
26 January 2003
,
W. Leonard
, mm ff; same as previous but
16 March 2003
, mm; same as previous but
23 March 2003
; same as previous but
22 December 2003
, mm ff;
Priest Point Park
,
Olympia
, elev. ca.
65 m
.
, ca.
47.07°N
, -
122.89°W
,
5 January 2003
,
W. Leonard
, mm f;
same as previous but
2 February 2003
, m; same as previous but
14 November 2004
, mm ff; along
Rock Candy Mtn. Road
,
Capitol State Forest
, elev. ca.
300 m
.
,
47.0256°N
, -
123.0735°W
,
18 December 2004
,
W. Leonard
, m.
Grays Harbor Co.
,
Porter Creek
Campground
,
Capitol State Forest
, elev.
90 m
.
,
46.9779°N
, -
123.2565°W
,
24 January 2005
,
W. Leonard
, m.
Cowlitz Co.
,
Germany
Creek Road
5.5 mi
N of Stella
at SR-4, elev.
110 m
.
,
46.2593°N
, -
123.1344°W
,
22 November 2003
,
W. Leonard
,
C. Richart
, mm; same as previous but
8 December 2003
, m f; same as previous but
11 November 2004
, mm ff.
Jefferson Co.
, along
Dosewallips River
,
Dosewallips Road
7.0 mi W of US-101,
Olympic National Forest
, elev.
115 m
.
,
47.7326°N
, -
123.0201°W
,
19 November 2003
,
W. Leonard
, m;
Falls View Campground
, US-101
7.8 mi
NW of Brinnon
, elev.
150 m
.
,
47.7893°N
, -
122.9255°W
,
22 February 2003
,
W. Leonard
, m f.
Mason Co.
, SR-119
6 mi
W of US-101 at
Hoodsport
, elev.
310 m
.
,
47.7326°N
, -
123.2097°W
,
7 February 2003
,
W. Leonard
, mm.
King Co.
, FR-9034 near
Granite
Mountain
Trailhead
,
Mt. Baker-Snoqualmie National Forest
, elev.
580 m
.
,
47.3977°N
, -
121.4855°W
,
W. Leonard
,
27 October 2007
, mm f; same as previous but
26 March 2004
,
W. Leonard
,
C. Richart
, m; same as previous but
25 October 2003
.
Lewis Co.
, 604
Roswell Road
,
Centralia
, elev.
65 m
.
,
46.7227°N
, -
122.9452°W
,
25 January 2004
,
C. Richart
,
W. Leonard
, mm ff;
SE end of
Riffle Lake
, elev.
250 m
.
,
46.4644°N
, -
122.1705°W
,
9 April 2004
,
W. Leonard
,
C. Richart
, m;
Slide Creek
,
Weyerhaeuser
4000
Rd
0.8 mi
SW of Pe-Ell-McDonald
Rd.
, elev.
190 m
.
,
46.5250°N
, -
123.1913°W
,
4 December 2004
,
W. Leonard
,
C. Richart
, mm ff; same as previous but
3 January 2005
,
W. Leonard
;
Stillman Basin
,
1.8 mi
on 4000
Weyerhaeuser
from
Pe-Ell
McDonald
Rd.
, elev.
290 m
.
,
46.5145°N
, -
123.2015°W
,
4 December 2004
,
W. Leonard
,
C. Richart
, from the litter and woody debris of a mixed forest including
Thuja plicata
,
Alnus rubra
,
Rubus spectabilus
,
Pseudotsuga menziesii
, and
Polystichum munitum
, m f;
North Fork of Newaukum River
, below confluence with
Middle Fork
, elev.
88 m
.
,
46.6046°N
, -
122.8481°W
,
29 December 2004
,
W. Leonard
, m.
Notes
:
This
species was described from
1 mi
W of Bayshore
and
4 mi
N of Shelton
,
Mason Co.
,
Washington
, and heretofore was known only from the
type
locality.
New
collecting revealed a much wider distribution, detailed above and in the supplemental KML map, and show this species to be the most widely distributed and most commonly collected conotylid in
western Washington. All
these populations are linked by identical anterior gonopods (
Fig. 192
), but the posterior gonopod coxites (
i.e.,
Fig. 193
) are so complicated that it is difficult to view them from exactly the same angle and make comparisons.
Thus
it is not possible really to be sure that all collections belong to the same species.
This
can be worked out later with genetic data.
Descriptions of new species of
Complicatella