A Review Of The Early Miocene Mastixioid Flora Of The Kristina Mine At Hrádek Nad Nisou In North Bohemia (The Czech Republic)
Author
Dedicated in memory of the late FrantiŠek Holý (1935 - 1984), an eminent Czech palaeobotanist
Author
Holý, František
National Museum, Václavské nám. 68,115 79 Prague 1, the Czech Republic
Author
Kvaček, Zlatko
Institute of Geology and Palaeontology, Faculty of Science, Charles University in Prague, Albertov 6, 128 43 Prague 2, the Czech Republic, e-mail: kvacek @ natur. cuni. cz
Author
Teodoridis, Vasilis
Department of Biology and Environmental Studies, Faculty of Education, Charles University in Prague, M. D. Rettigové 4, 116 39 Prague 1, the Czech Republic, e-mail: vasilis. teodoridis @ pedf. cuni. cz
text
Acta Musei Nationalis Pragae Series B
2012
2012-12-31
68
3 - 4
53
118
journal article
10.5281/zenodo.13191145
2533-4069
13191145
Laurophyllum pseudoprinceps
WEYLAND
et
KILPPER
Pl. 1, figs 10, 12-14, pl. 11, figs 2-3
1963
1963
1963
1971
Laurophyllum pseudoprinceps
WEYLAND
et
KILPPER
, p. 100, pl. 23, figs 14-19, text-fig. 6 (Frimmersdorf Mine, Ville).
Laurophyllum verrucosum
WEYLAND
et
KILPPER
, p. 102, pl. 24, figs 24-25, pl. 25, figs 26-27 (Frimmersdorf Mine, Ville).
Laurophyllum undulatum
WEYLAND
et
KILPPER
, p. 101, pl. 24, figs 20-21, text-fig. 7-8 (Frimmersdorf Mine, Ville).
Laurophyllum pseudoprinceps
WEYLAND
et
KILPPER
; Kvaček, p. 50, pl. 1, figs 4-6, pl. 3, figs 3-5, pl. 4, figs 1-4, text-fig. 1 (Hrádek/N., Kristina Mine, many other sites in North Bohemian Oligocene and Miocene).
Incomplete and fragmentary simple leaves, lamina elliptic to ovate,
37–67 mm
long,
12–27 mm
wide, base cuneate, decurrent into an up to
13 mm
long and
2 mm
wide petiole, apex acuminate and blunt, margin entire, venation brochidodromous, midrib strong, straight or slightly curved in apical part, secondary veins thinner, straight, looping by margin, alternate, originating at an angle of 30–50°, tertiary veins perpendicular, straight to sinuous, venation of the higher orders regular polygonal reticulate, areolation well developed, 4-sided, veinlets dichotomous to dendritic branching. Adaxial epidermis thickly cutinized, partly granulate or smooth, non-modified cells polygonal, anticlinal walls varying from almost straight to strongly undulate, mostly exhibiting bead-like thickening. Abaxial cuticle usually also strongly cutinized, non-modified cells showing polygonal to strongly lobately wavy outlines, stomata amphibrachyparacytic, usually 17–25 µm long and 15–20 µm wide, outer subsidiary cells partly unequally developed. Trichome bases simple, sporadic. Lens-shaped oil cells common.
D i s c u s s i o n: This element is very common in the Miocene and Oligocene of Europe but seems to be endemic to Europe. Due to autecological variation (sun vs shade leaves) extreme forms have usually been assigned to independent fossil species (
Weyland and Kilpper 1963
) or even genera (
Juchniewicz 1975
). We support the previous interpretations by
Kvaček (1971)
and others and consider them as ecotypes of the same species. The stomatal
type
suggests an affinity to
Ocotea
(hence not generally accepted
Ocoteophyllum
JUCHNIEWICZ
1975
or the direct transfer to
Ocotea
by
Uzunova and Stojanova 1999
) but no exact living counterpart in today’s natural world has been suggest- ed so far to match in all respects. Therefore we hesitate to accept the direct transfer into this living genus (see also
Kvaček et al. 2011
).
M a t e r i a l s t u d i e d: Isolated leaf compressions on slides, G 8894-8961 (KR 31, 32, 36, 51, 53, 61, 62, 63, 85, 124, 125, 127, 131, 144, 146, 149, 150, 161, 163, 174, 189, 191, 227, 228, 230, 231, 263, 296, 297, 298, 317, 321, 322, 408, 409, 528, sine num.).