The Nemourinae (Insecta, Nemouridae) of the eastern Nearctic Author Grubbs, Scott A. 0000-0003-2170-9716 Department of Biology and Center for Biodiversity Studies, Western Kentucky University, Bowling Green, Kentucky, 42101, USA Department of Biology and Monte L. Bean Life Science Museum, Brigham Young University, Provo, Utah, 84602, USA & scott. grubbs @ wku. edu; https: // orcid. org / 0000 - 0003 - 2170 - 9716 scott.grubbs@wku.edu Author Baumann, Richard W. 0000-0003-0838-4080 richard _ baumann @ byu. edu; https: // orcid. org / 0000 - 0003 - 0838 - 4080 richard_baumann@byu.edu text Zootaxa 2023 2023-06-19 5306 1 1 53 http://dx.doi.org/10.11646/zootaxa.5306.1.1 journal article 10.11646/zootaxa.5306.1.1 1175-5326 8054287 21AD4F85-E313-47CB-BA98-9896D2B1DC05 22. Soyedina vallicularia ( Wu, 1923 ) Valley Forestfly http://lsid.speciesfile.org/urn:lsid: Plecoptera .speciesfile.org:TaxonName:6084 ( Figs. 145–150 ) Nemoura vallicularia Wu 1923:58 . Holotype male ( Cornell University Insect Collection ), Ithaca , ( Tompkins Co. ), New York , USA Nemoura vallicularia Needham & Claassen, 1925:215 Nemoura (Soyedina) vallicularia Ricker, 1952:53 Soyedina vallicularia Illies, 1966:248 Soyedina vallicularia Zwick, 1973:357 Nemoura (Soyedina) vallicularia Hitchcock, 1974:110 Soyedina vallicularia Baumann, 1975:67 Soyedina vallicularia Stewart & Stark, 1988:188 Soyedina vallicularia Stewart & Stark, 2002:215 Soyedina vallicularia Grubbs, 2006:40 Soyedina vallicularia Stewart & Stark, 2002:215 Soyedina vallicularia Stark, 2017:215 Soyedina vallicularia Grubbs & Baumann, 2019:227 Distribution. Canada : LB, NS, ON, PQ. USA ; CT, IA, IL, IN, KY, MA, MD, ME, MI, NY, OH, PA, TN, VA, WI, WV ( DeWalt et al. 2022 ) Male . Macropterous. Body length 4.7–5.1 mm , forewing length 6.8–8.1 mm (n = 16). Gills absent. Cerci simple and unmodified. Paraprocts with two lobes, inner lobes lightly sclerotized and inconspicuous, outer lobes sclerotized, in basal half bulbous, tapered and straight-sided in apical half ( Fig. 149 ). Epiproct asymmetric, typical for genus ( Fig. 145 ); ventral sclerites partially recurved over abdomen, distinctly asymmetric, apically narrowly triangular ( Fig. 145 ); dorsal sclerites distinctly asymmetric, appearing scaly at higher magnifications ( Figs. 145‒ 148 ); open apically, inner member heavily sclerotized, flanged apically, lip asymmetric with apical half longer than basal portion ( Figs. 145‒146 ). Female . Macropterous. Body length 5.7–8.2 mm , forewing length 7.8.– 9.7 mm (n = 20). Gills absent. Cerci simple and unmodified. Subgenital plate well-developed, subtriangular in shape; base broad, extending from posterior of seventh sternum entirely over eighth sternum and approximately ¼ over ninth sternum; terminating in broadly-rounded apex ( Fig. 150 ). Larva. Described in full (see Stewart & Stark 2002 and all references therein). Comments. Soyedina vallicularia is the most widespread amongst the eastern Nearctic Soyedina , including the Appalachian Mountains , Great Lakes region , and Ohio River and upper Mississippi River basins ( Grubbs & Baumann 2019 ) .